Ecological Traitdata List

This is an exemplary list of traits following the Ecological Traitdata Standard (ETS), version 0.4

The ETS (https://ecologicaltraitdata.github.io/ETS/) defines a vocabulary of essential columns for raw data of functional traits for ecological research.

The ETS is organized into a core section with essential columns for trait data, extensions which are allowing to provide additional layers of information, as well as a vocabulary for metadata information of particular importance for trait data. A final section provides defined terms for lists of trait definitions, also termed a Trait Thesaurus.

The following list of traits serve as an example of a Trait Thesaurus. It is based around traits for invertebrates, but definitions were made with potential applicability to other animal groups. Nevertheless, this Trait Thesaurus is not an attempt at a complete Thesaurus for all animal groups, but rather a starting point for other Thesauri.

Structure of the trait data:

Each trait is assigned to one of seven trait classes, which differentiate broad types of traits. Traits with very general definitions form the highest level of the trait hierarchy, with more specific traits being assigned to those general traits. Each traits’ position in the trait hierarchy is defined through the broaderTerm column, which indicates the next higher-level trait. Any trait can be defined as the broaderTerm for more than one trait. Lowest-level traits are often a part of a measurement or range, e.g. maximum, minimum, average values.

For example: Clutch_size_max is a more specific definition of Clutch_size which itself is a specific aspect of Reproduction_strategy which is a specific description of Reproduction_mode which is one of the high-level traits in the class life-history. The high-level trait Reproduction_mode is defined as the broaderTerm for seven traits.

Categorical traits are defined by factor levels, which is in contrast to other Trait Thesauri where each factor level is an individual trait of boolean class. In this Trait Thesaurus, factor levels are suggestions without being intended to be exhaustive.

Numerical traits include information on the traitUnit, which is essential for complementarity with other trait databases, and information on maxAllowed and minAllowed values. Those value ranges encompass the range of mathematically possible values (e.g. Clutch_size cannot be negative) but does not reflect a biologically meaningful range.

For each trait, a source is given and its relation to that source is indicated in relationSource:

• equals if the trait is taken directly from the source without modification

• takenfrom if the trait is based on a trait list which does not include formal descriptions or trait-specific references (e.g. the trait was used in a scientific publication or is provided in taxonomic literature)

• refines if the trait gives a more precice definition of the original trait

• broadens if the trait gives a more general definition of the original trait

• inheritsfrom if the trait is slight modification of the original trait but changes in the definition of the original trait should be reflected in this trait as well (e.g. Body_length_max is a modification of Body_length).

Defined traits:

behaviour
Protection_behaviour | Water_regulation_behaviour | Thermal_regulation_behaviour | Phenology | Activity_period | Observation_first_date | Observation_last_date | Activity_period_length | Flight_period_length | Dispersal_mode | Dispersal_effectiveness | Dispersal_range_active | Dispersal_range_passive | Flight_ability | Intraspecific_relation | Clustering_of_individuals | Social_strategy | Social_structure | Number_queens | Protection_behaviour | Hibernation_strategy | Hibernation_in_conifers | Reproduction_behaviour | Physical_eggs_protection | Egg_protection_type | Nest_stratum | Egg_placement | Pupal_location | Parental_care |
environment
Altitude | Altitude_max | Altitude_min | Altitude_class | Habitat_specialization | Habitat_preferences | Ecological_niche | Micro_habitat | Humidity | Humidity_soil | Humidity_soil_max | Humidity_soil_min | Humidity_air | Humidity_air_max | Humidity_air_min | Oxygen_water | Oxygen_water_max | Temperature_survival | Temperature_survival_max | Temperature_survival_min | Temperature_activity | Temperature_activity_max | Temperature_activity_min | Temperature_reproduction | Temperature_reproduction_max | Temperature_reproduction_min | Temperature_development | Temperature_development_max | Temperature_development_min | Stratum_preference | Stratum_phenology | Habitat_CORINE | Habitat_forest | Habitat_grassland | Nature_Orientation | Climate_zones_Köppen | Relict_species | Oxygen_water_min | Deadwood_dependent |
feeding
Stable_Isotopes | Feeding_guild | Feeding_guild_adults | Feeding_guild_juveniles | Feeding_mode | Feeding_mode_juveniles | Feeding_specialization | Host | Host_number | Endoghagous_lifestile | Feeding_specialization_juveniles | Feeding_tissue | Feeding_plant_part | Feeding_plant_part_juveniles | Feeding_tissue_juveniles | Feeding_behaviour | Trap_strategy | Trophic_level | Trophic_level_num |
life-history
Sexuality | Asexual_strategy | Reproduction_mode | Reproduction_strategy | Clutch_size | Clutch_size_max | Clutch_size_min | Nr_eggs_lifetime | Nr_eggs_lifetime_max | Nr_eggs_lifetime_min | Egg_size | Egg_length | Egg_length_max | Egg_length_min | Egg_width | Egg_width_max | Egg_width_min | Egg_diameter | Egg_diameter_max | Egg_diameter_min | Egg_volume | Egg_volume_max | Egg_volume_min | Egg_weight | Egg_weight_max | Egg_weight_min | Egg_biomass | Egg_development_time | Egg_development_time_max | Egg_development_time_min | Developmental_stages | Developmental_stages_number | Nr_larval_stages | Developmental_stages_time | Developmental_stages_time_max | Developmental_stages_time_min | Fecundity | Generation_number_year | Generation_number_year_max | Generation_number_year_min | Reproduction_events_year | Reproduction_events_year_max | Reproduction_events_year_min | Reproduction_events_life | Reproduction_events_life_max | Reproduction_events_life_min | Mating_system | Life_span_potential | Life_span_potential_max | Life_span_potential_min | Life_span_observed | Life_span_observed_max | Life_span_observed_min | Development | Age_at_maturity | Age_at_maturity_max | Age_at_maturity_min | Age_at_reproduction | Age_at_reproduction_max | Age_at_reproduction_min |
morphology
Reproduction_morphology | Sperm_storage | Nr_Ovariols | Nr_Ovariols_max | Nr_Ovariols_min | Body_shape | Body_size | Body_length | Body_length_max | Body_length_min | Pronotum_length | Pronotum_length_max | Pronotum_length_min | Thorax_length | Thorax_length_max | Thorax_length_min | Head_length | Head_length_max | Head_length_min | Body_width | Body_width_max | Body_width_min | Pronotum_width | Pronotum_width_max | Pronotum_width_min | Intertegular_span | Intertegular_span_max | Intertegular_span_min | Thorax_width | Thorax_width_max | Thorax_width_min | Head_width | Head_width_max | Head_width_min | Body_height | Body_height_max | Body_height_min | Pronotum_height | Pronotum_height_max | Pronotum_height_min | Thorax_height | Thorax_height_max | Thorax_height_min | Head_height | Head_height_max | Head_height_min | Body_mass_live | Body_mass_live_max | Body_mass_live_min | Body_mass_fresh | Body_mass_fresh_max | Body_mass_fresh_min | Body_mass_dried | Body_mass_dried_max | Body_mass_dried_min | Body_volume | Body_volume_max | Body_volume_min | Length_width_ratio | Length_width_ratio_max | Length_width_ratio_min | Length_width_ratio_pronotum | Length_width_ratio_pronotum_max | Length_width_ratio_pronotum_min | Length_width_ratio_thorax | Length_width_ratio_thorax_max | Length_width_ratio_thorax_min | Length_width_ratio_head | Length_width_ratio_head_max | Length_width_ratio_head_min | Nutrition_morphology | Mouthpart_type | Mandible_width | Mandible_width_max | Mandible_width_min | Mandible_length | Mandible_length_max | Mandible_length_min | Tongue_length | Tongue_length_max | Tongue_length_min | Rostrum_length | Rostrum_length_max | Rostrum_length_min | Rostrum_width | Rostrum_width_max | Rostrum_width_min | Rostrum_ratio | Rostrum_ratio_max | Rostrum_ratio_min | Lipidic_storage | Respiratory_organ_type | Sensory_morphology | Antenna_length | Antenna_length_max | Antenna_length_min | Antenna_segment1_length | Antenna_segment2_length | Antenna_segment3_length | Antenna_segment4_length | Antenna_segment5_length | Antenna_segment6_length | General_Body_pubescence | Chemoreceptor | Thermoreceptor | Mechanoreceptor | Photoreceptor | Eye_number | Eye_area | Eye_width | Eye_width_max | Eye_width_min | Eye_diameter | Eye_diameter_max | Eye_diameter_min | Locomotion_morphology | Leg_length | Leg_length_max | Leg_length_min | Hind_leg_length | Hind_leg_length_max | Hind_leg_length_min | Mid_leg_length | Mid_leg_length_max | Mid_leg_length_min | Front_leg_length | Front_leg_length_max | Front_leg_length_min | Femur_length | Femur_length_max | Femur_length_min | Front_femur_length | Front_femur_length_max | Front_femur_length_min | Mid_femur_length | Mid_femur_length_max | Mid_femur_length_min | Hind_femur_length | Hind_femur_length_max | Hind_femur_length_min | Tibia_length | Tibia_length_max | Tibia_length_min | Front_tibia_length | Front_tibia_length_max | Front_tibia_length_min | Mid_Tibia_length | Mid_Tibia_length_max | Mid_Tibia_length_min | Hind_tibia_length | Hind_tibia_length_max | Hind_tibia_length_min | Trochanter_length | Trochanter_length_max | Trochanter_length_min | Front_trochanter_length | Front_trochanter_length_max | Front_trochanter_length_min | Mid_trochanter_length | Mid_trochanter_length_max | Mid_trochanter_length_min | Hind_trochanter_length | Hind_trochanter_length_max | Hind_trochanter_length_min | Femur_width | Femur_width_max | Femur_width_min | Front_femur_width | Front_femur_width_max | Front_femur_width_min | Mid_femur_width | Mid_femur_width_max | Mid_femur_width_min | Hind_femur_width | Hind_femur_width_max | Hind_femur_width_min | Wing_appearance | Wing_size_relation | Front_wing_width | Front_wing_width_max | Front_wing_width_min | Front_wing_surface | Front_wing_surface_max | Front_wing_surface_min | Hind_wing_length | Hind_wing_length_max | Hind_wing_length_min | Hind_wing_width | Hind_wing_width_max | Hind_wing_width_min | Hind_wing_surface | Hind_wing_surface_max | Hind_wing_surface_min | Wing_appearance_Females | Wing_appearance_Males | Front_wing_length | Front_wing_length_max | Front_wing_length_min | Elytra_length | Elytra_length_max | Elytra_length_min | Elytra_width | Elytra_width_max | Elytra_width_min | Elytra_surface | Elytra_surface_max | Elytra_surface_min | Radial_nervure_length | Radial_nervure_length_max | Radial_nervure_length_min | Median_nervure_length | Median_nervure_length_max | Median_nervure_length_min | Front_wing_span | Front_wing_span_max | Front_wing_span_min | Hind_wing_span | Hind_wing_span_max | Hind_wing_span_min | Saltatory_apparatus | Saltatory_apparatus_length | Saltatory_apparatus_length_max | Saltatory_apparatus_length_min | Leg_body_ratio | Leg_body_ratio_max | Leg_body_ratio_min | Protection_morphology | Body_color | Body_color_gradient | Body_color_pattern | Repellent_glands | Sclerotization | Detoxication_strategy | Water_regulation_morphology |
physiology
Water_regulation_physiology | Thermal_regulation_physiology | Water_loss_rate | Larval_inundation_tolerance | Venom_excretion | Regeneration_strategy |
pollination
Flower_color_preference | Flower_visitor | Flower_specialisation | Flower_feeding |

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Traits on behaviour

Protection_behaviour

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	Protection_behaviour
traitName	Protection_behaviour
trait_class	behaviour
broaderTerm	https://ecologicaltraitdata.github.io/TraitDataList/#behaviour
valueType	factor
traitUnit	NA
maxAllowedValue	NA
minAllowedValue	NA
factorLevels	hibernation/starvation
traitDescription	strategies used for protection
Comments	Any animal behaviour allowing it to escape from a danger or a nocive situation and to survive
example	hibernation
relationSource	equals
source	http://t-sita.cesab.org/BETSI_vizInfo.jsp?trait=Protection_behaviour

Water_regulation_behaviour

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	Water_regulation_behaviour
traitName	Water_regulation_behaviour
trait_class	behaviour
broaderTerm	https://ecologicaltraitdata.github.io/TraitDataList/#behaviour
valueType	character
traitUnit	NA
maxAllowedValue	NA
minAllowedValue	NA
factorLevels	NA
traitDescription	any behavioural adaptation which ensures an hydric status of the animal
Comments
example	NA
relationSource	refines
source	http://t-sita.cesab.org/BETSI_vizInfo.jsp?trait=waterregulation%20strategies

Thermal_regulation_behaviour

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	Thermal_regulation_behaviour
traitName	Thermal_regulation_behaviour
trait_class	behaviour
broaderTerm	https://ecologicaltraitdata.github.io/TraitDataList/#behaviour
valueType	character
traitUnit	NA
maxAllowedValue	NA
minAllowedValue	NA
factorLevels	NA
traitDescription	any behavioural adaptation which keeps the body temperature within certain boundaries
Comments
example	NA
relationSource	refines
source	http://t-sita.cesab.org/BETSI_vizInfo.jsp?trait=Thermoregulation_strategies

Phenology

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	Phenology
traitName	Phenology
trait_class	behaviour
broaderTerm	https://ecologicaltraitdata.github.io/TraitDataList/#behaviour
valueType	character
traitUnit	NA
maxAllowedValue	NA
minAllowedValue	NA
factorLevels	NA
traitDescription	any behavioural adaptation which manifests during specific times
Comments
example	NA
relationSource	equals
source	http://t-sita.cesab.org/BETSI_vizInfo.jsp?trait=Phenology

Activity_period

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	Activity_period
traitName	Activity_period
trait_class	behaviour
broaderTerm	https://ecologicaltraitdata.github.io/TraitDataList/#Phenology
valueType	character
traitUnit	NA
maxAllowedValue	NA
minAllowedValue	NA
factorLevels	NA
traitDescription	first and last month of a year when individuals can be observed
Comments	First and last month are abbreviated to three letters based on english names. Months are connected with “-” for continous time spans or separated with “.” for interupted time spans.
example	Apr-Aug.Nov
relationSource	refines
source	http://t-sita.cesab.org/BETSI_vizInfo.jsp?trait=Adult_activity

Observation_first_date

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	Observation_first_date
traitName	Observation_first_date
trait_class	behaviour
broaderTerm	https://ecologicaltraitdata.github.io/TraitDataList/#Activity_period
valueType	integer
traitUnit	yearday
maxAllowedValue	365
minAllowedValue	1
factorLevels	NA
traitDescription	earliest observation within a year, given as yearday
Comments
example	125
relationSource	refines
source	http://t-sita.cesab.org/BETSI_vizInfo.jsp?trait=Adult_activity

Observation_last_date

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	Observation_last_date
traitName	Observation_last_date
trait_class	behaviour
broaderTerm	https://ecologicaltraitdata.github.io/TraitDataList/#Activity_period
valueType	integer
traitUnit	yearday
maxAllowedValue	365
minAllowedValue	1
factorLevels	NA
traitDescription	latest observation within a year, given as yearday
Comments
example	350
relationSource	refines
source	http://t-sita.cesab.org/BETSI_vizInfo.jsp?trait=Adult_activity

Activity_period_length

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	Activity_period_length
traitName	Activity_period_length
trait_class	behaviour
broaderTerm	https://ecologicaltraitdata.github.io/TraitDataList/#Phenology
valueType	integer
traitUnit	month
maxAllowedValue	12
minAllowedValue	0
factorLevels	NA
traitDescription	number of months during individuals are active
Comments	Weeks and days are expressed as fractions of an an average month (i.e. 30 days)
example	3
relationSource	refines
source	http://t-sita.cesab.org/BETSI_vizInfo.jsp?trait=Adult_activity

Flight_period_length

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	Flight_period_length
traitName	Flight_period_length
trait_class	behaviour
broaderTerm	https://ecologicaltraitdata.github.io/TraitDataList/#Activity_period_length
valueType	integer
traitUnit	month
maxAllowedValue	12
minAllowedValue	0
factorLevels	NA
traitDescription	number of months during which flying individuals are observed
Comments	Weeks and days are expressed as fractions of an an average month (i.e. 30 days)
example	1.5
relationSource	refines
source	http://t-sita.cesab.org/BETSI_vizInfo.jsp?trait=Adult_activity

Dispersal_mode

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	Dispersal_mode
traitName	Dispersal_mode
trait_class	behaviour
broaderTerm	https://ecologicaltraitdata.github.io/TraitDataList/#behaviour
valueType	factor
traitUnit	NA
maxAllowedValue	NA
minAllowedValue	NA
factorLevels	active/passive/both
traitDescription	strategies used for dispersal
Comments	Does not include disperal of offspring or reproductive cells
example	active
relationSource	equals
source	http://t-sita.cesab.org/BETSI_vizInfo.jsp?trait=Dispersal_behaviour

Dispersal_effectiveness

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	Dispersal_effectiveness
traitName	Dispersal_effectiveness
trait_class	behaviour
broaderTerm	https://ecologicaltraitdata.github.io/TraitDataList/#Dispersal_mode
valueType	factor
traitUnit	NA
maxAllowedValue	NA
minAllowedValue	NA
factorLevels	0/0.25/0.5/0.75/1
traitDescription	dispersal ability relative to other species within the same taxonomic group
Comments
example	0.5
relationSource	takenfrom
source	https://www.nature.com/articles/sdata201513

Dispersal_range_active

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	Dispersal_range_active
traitName	Dispersal_range_active
trait_class	behaviour
broaderTerm	https://ecologicaltraitdata.github.io/TraitDataList/#Dispersal_mode
valueType	factor
traitUnit	NA
maxAllowedValue	NA
minAllowedValue	NA
factorLevels	meters/kilometers/10kilometers/100kilometers/continents/global
traitDescription	potential range of dispersal using active dispersal strategies
Comments
example	10kilometers
relationSource	refines
source	http://t-sita.cesab.org/BETSI_vizInfo.jsp?trait=Dispersal_behaviour

Dispersal_range_passive

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	Dispersal_range_passive
traitName	Dispersal_range_passive
trait_class	behaviour
broaderTerm	https://ecologicaltraitdata.github.io/TraitDataList/#Dispersal_mode
valueType	factor
traitUnit	NA
maxAllowedValue	NA
minAllowedValue	NA
factorLevels	meters/kilometers/10kilometers/100kilometers/continents/global
traitDescription	potential range of dispersal using passive dispersal strategies
Comments
example	10kilometers
relationSource	refines
source	http://t-sita.cesab.org/BETSI_vizInfo.jsp?trait=Dispersal_behaviour

Flight_ability

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	Flight_ability
traitName	Flight_ability
trait_class	behaviour
broaderTerm	https://ecologicaltraitdata.github.io/TraitDataList/#Dispersal_mode
valueType	logical
traitUnit	NA
maxAllowedValue	NA
minAllowedValue	NA
factorLevels	NA
traitDescription	locomotion by means of wings
Comments
example	TRUE
relationSource	refines
source	http://t-sita.cesab.org/BETSI_vizInfo.jsp?trait=Dispersal_behaviour

Intraspecific_relation

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	Intraspecific_relation
traitName	Intraspecific_relation
trait_class	behaviour
broaderTerm	https://ecologicaltraitdata.github.io/TraitDataList/#behaviour
valueType	character
traitUnit	NA
maxAllowedValue	NA
minAllowedValue	NA
factorLevels	NA
traitDescription	any behavioural adaptation which increases interactions within the species
Comments
example	NA
relationSource	equals
source	http://t-sita.cesab.org/BETSI_vizInfo.jsp?trait=Intraspecific_relation

Clustering_of_individuals

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	Clustering_of_individuals
traitName	Clustering_of_individuals
trait_class	behaviour
broaderTerm	https://ecologicaltraitdata.github.io/TraitDataList/#Intraspecific_relation
valueType	logical
traitUnit	NA
maxAllowedValue	NA
minAllowedValue	NA
factorLevels	NA
traitDescription	individuals usually occur in spatial clusters
Comments
example	TRUE
relationSource
source

Social_strategy

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	Social_strategy
traitName	Social_strategy
trait_class	behaviour
broaderTerm	https://ecologicaltraitdata.github.io/TraitDataList/#Intraspecific_relation
valueType	factor
traitUnit	NA
maxAllowedValue	NA
minAllowedValue	NA
factorLevels	subsocial/solitary_but_social/communal/quasisocial/semisocial/eusocial
traitDescription	degree to which individuals in an animal population tend to associate in social groups and form cooperative societies
Comments	Any behavioural or ecological features that increase interindividual attraction, and tolerance.
example	eusocial
relationSource	refines
source	http://t-sita.cesab.org/BETSI_vizInfo.jsp?trait=Sociability

Social_structure

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	Social_structure
traitName	Social_structure
trait_class	behaviour
broaderTerm	https://ecologicaltraitdata.github.io/TraitDataList/#Social_strategy
valueType	character
traitUnit	NA
maxAllowedValue	NA
minAllowedValue	NA
factorLevels	NA
traitDescription	description of the social structure
Comments	details wether cooperation includes feeding, reproduction or protection
example	NA
relationSource	refines
source	http://t-sita.cesab.org/BETSI_vizInfo.jsp?trait=Sociability

Number_queens

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	Number_queens
traitName	Number_queens
trait_class	behaviour
broaderTerm	https://ecologicaltraitdata.github.io/TraitDataList/#Social_structure
valueType	numeric
traitUnit	count
maxAllowedValue	999
minAllowedValue	0
factorLevels	NA
traitDescription	number of reproducing females within one colony/state/hive
Comments
example	2
relationSource	refines
source	http://t-sita.cesab.org/BETSI_vizInfo.jsp?trait=Sociability

Protection_behaviour

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	Protection_behaviour
traitName	Protection_behaviour
trait_class	behaviour
broaderTerm	https://ecologicaltraitdata.github.io/TraitDataList/#behaviour
valueType	factor
traitUnit	NA
maxAllowedValue	NA
minAllowedValue	NA
factorLevels	hibernation/starvation
traitDescription	strategies used for protection
Comments	Any animal behaviour allowing it to escape from a danger or a nocive situation and to survive
example	hibernation
relationSource	equals
source	http://t-sita.cesab.org/BETSI_vizInfo.jsp?trait=Protection_behaviour

Hibernation_strategy

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	Hibernation_strategy
traitName	Hibernation_strategy
trait_class	behaviour
broaderTerm	https://ecologicaltraitdata.github.io/TraitDataList/#Protection_behaviour
valueType	factor
traitUnit	NA
maxAllowedValue	NA
minAllowedValue	NA
factorLevels	dormancy/resting/freezing
traitDescription	strategy against climatic adversity, such as low temperature or drought
Comments
example	dormancy
relationSource	equals
source	http://t-sita.cesab.org/BETSI_vizInfo.jsp?trait=Overwintering

Hibernation_in_conifers

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	Hibernation_in_conifers
traitName	Hibernation_in_conifers
trait_class	behaviour
broaderTerm	https://ecologicaltraitdata.github.io/TraitDataList/#Hibernation_strategy
valueType	logical
traitUnit	NA
maxAllowedValue	NA
minAllowedValue	NA
factorLevels	NA
traitDescription	coniferous tree species are used for hibernation
Comments
example	TRUE
relationSource	refines
source	https://ecologicaltraitdata.github.io/TraitDataList/#Hibernation_strategy

Reproduction_behaviour

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	Reproduction_behaviour
traitName	Reproduction_behaviour
trait_class	behaviour
broaderTerm	https://ecologicaltraitdata.github.io/TraitDataList/#behaviour
valueType	character
traitUnit	NA
maxAllowedValue	NA
minAllowedValue	NA
factorLevels	NA
traitDescription	any behavioural adaptation which increases the odds of succesful reproduction
Comments
example	NA
relationSource	equals
source	http://t-sita.cesab.org/BETSI_vizInfo.jsp?trait=Reproduction_behaviour

Physical_eggs_protection

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	Physical_eggs_protection
traitName	Physical_eggs_protection
trait_class	behaviour
broaderTerm	https://ecologicaltraitdata.github.io/TraitDataList/#Reproduction_behaviour
valueType	logical
traitUnit	NA
maxAllowedValue	NA
minAllowedValue	NA
factorLevels	NA
traitDescription	protection of eggs by physical means
Comments	Includes protection outside the parent and inside the parend
example	TRUE
relationSource	equals
source	http://t-sita.cesab.org/BETSI_vizInfo.jsp?trait=Physical_eggs_protection

Egg_protection_type

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	Egg_protection_type
traitName	Egg_protection_type
trait_class	behaviour
broaderTerm	https://ecologicaltraitdata.github.io/TraitDataList/#Physical_eggs_protection
valueType	factor
traitUnit	NA
maxAllowedValue	NA
minAllowedValue	NA
factorLevels	nest/internal/parasitism
traitDescription	strategies for egg protection
Comments	Parasitims includes cuckoldry. Internal protection applies to viviparous strategies
example	internal
relationSource	refines
source	https://ecologicaltraitdata.github.io/TraitDataList/#Egg_protection_type

Nest_stratum

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	Nest_stratum
traitName	Nest_stratum
trait_class	behaviour
broaderTerm	https://ecologicaltraitdata.github.io/TraitDataList/#Egg_protection_type
valueType	factor
traitUnit	NA
maxAllowedValue	NA
minAllowedValue	NA
factorLevels	burrow/ground/vegetation
traitDescription	stratum of the nest location
Comments	Vegetation includes low and dead vegetation
example	ground
relationSource	refines
source	https://ecologicaltraitdata.github.io/TraitDataList/#Egg_protection_type

Egg_placement

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	Egg_placement
traitName	Egg_placement
trait_class	behaviour
broaderTerm	https://ecologicaltraitdata.github.io/TraitDataList/#Egg_protection_type
valueType	factor
traitUnit	NA
maxAllowedValue	NA
minAllowedValue	NA
factorLevels	soil/ground/vegetation/stem/leaf/leaf_underside/leaf_upside/bark/wood/living_host/sedated_host/carcass/dung
traitDescription	microhabitat in which eggs are placed
Comments	defined when eggs are placed without construction of additional supporting structures
example	ground
relationSource	refines
source	https://ecologicaltraitdata.github.io/TraitDataList/#Egg_protection_type

Pupal_location

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	Pupal_location
traitName	Pupal_location
trait_class	behaviour
broaderTerm	https://ecologicaltraitdata.github.io/TraitDataList/#Reproduction_behaviour
valueType	character
traitUnit	NA
maxAllowedValue	NA
minAllowedValue	NA
factorLevels	NA
traitDescription	position of the pupa within the nest
Comments	Usually applies to burrows or tree holes
example	NA
relationSource	refines
source	https://ecologicaltraitdata.github.io/TraitDataList/#Egg_protection_type

Parental_care

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	Parental_care
traitName	Parental_care
trait_class	behaviour
broaderTerm	https://ecologicaltraitdata.github.io/TraitDataList/#Reproduction_behaviour
valueType	factor
traitUnit	NA
maxAllowedValue	NA
minAllowedValue	NA
factorLevels	females/males/both_sexes/workers/none
traitDescription	investment in protection or nurturing of offspring
Comments
example	females
relationSource	refines
source	http://t-sita.cesab.org/BETSI_vizInfo.jsp?trait=Parental_care

Traits relating to the environment

Altitude

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	Altitude
traitName	Altitude
trait_class	environment
broaderTerm	https://ecologicaltraitdata.github.io/TraitDataList/#environment
valueType	numeric
traitUnit	m
maxAllowedValue	9999
minAllowedValue	0
factorLevels	NA
traitDescription	average elevation above sea level at which individuals have been observed
Comments
example	250
relationSource	refines
source	http://t-sita.cesab.org/BETSI_vizInfo.jsp?trait=Altitude

Altitude_max

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	Altitude_max
traitName	Altitude_max
trait_class	environment
broaderTerm	https://ecologicaltraitdata.github.io/TraitDataList/#Altitude
valueType	numeric
traitUnit	m
maxAllowedValue	9999
minAllowedValue	0
factorLevels	NA
traitDescription	highest elevation above sea level at which individuals have been observed
Comments
example	250
relationSource	refines
source	http://t-sita.cesab.org/BETSI_vizInfo.jsp?trait=Altitude

Altitude_min

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	Altitude_min
traitName	Altitude_min
trait_class	environment
broaderTerm	https://ecologicaltraitdata.github.io/TraitDataList/#Altitude
valueType	numeric
traitUnit	m
maxAllowedValue	9999
minAllowedValue	0
factorLevels	NA
traitDescription	lowest elevation above sea level at which individuals have been observed
Comments
example	250
relationSource	refines
source	http://t-sita.cesab.org/BETSI_vizInfo.jsp?trait=Altitude

Altitude_class

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	Altitude_class
traitName	Altitude_class
trait_class	environment
broaderTerm	https://ecologicaltraitdata.github.io/TraitDataList/#Altitude
valueType	factor
traitUnit	NA
maxAllowedValue	NA
minAllowedValue	NA
factorLevels	planar/kollin/montan/subalpin/alpin/nival
traitDescription	occurence of individuals is restricted to a certain altitude class
Comments	planar: to 400m/kollin: 400-800m/montan: 800-1.500/subalpin: 1.500-2.300/alpin: 2.300-2.700/nival: above 2.700
example	alpin
relationSource	refines
source	http://t-sita.cesab.org/BETSI_vizInfo.jsp?trait=Altitude

Habitat_specialization

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	Habitat_specialization
traitName	Habitat_specialization
trait_class	environment
broaderTerm	https://ecologicaltraitdata.github.io/TraitDataList/#environment
valueType	logical
traitUnit	NA
maxAllowedValue	NA
minAllowedValue	NA
factorLevels	NA
traitDescription	occurence of individuals is restricted to a certain habitat
Comments	Any level of habitat specialization should be indicated here.
example	TRUE
relationSource	refines
source	http://t-sita.cesab.org/BETSI_vizInfo.jsp?trait=Habitat

Habitat_preferences

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	Habitat_preferences
traitName	Habitat_preferences
trait_class	environment
broaderTerm	https://ecologicaltraitdata.github.io/TraitDataList/#Habitat_specialization
valueType	integer
traitUnit	NA
maxAllowedValue	9999
minAllowedValue	0
factorLevels	NA
traitDescription	number of habitat types where individuals can be observed
Comments	Obiquitos species are assigned 0 and strict specialists are assigned 1.
example	2
relationSource	refines
source	http://t-sita.cesab.org/BETSI_vizInfo.jsp?trait=Habitat

Ecological_niche

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	Ecological_niche
traitName	Ecological_niche
trait_class	environment
broaderTerm	https://ecologicaltraitdata.github.io/TraitDataList/#Habitat_preferences
valueType	factor
traitUnit	NA
maxAllowedValue	NA
minAllowedValue	NA
factorLevels	azidophil/chromophil/coprophil/halophil/heliophil/hygrophil/hylophil/krenophil/mycetophil/myrmecophil/necrophil/osmophil/petrophil/pholeophil/psammophil/rheophil/saprophil/silicophil/thermophil/tixophil/troglophil/tyrophil/xerophil
traitDescription	species distribution is defined by an environmental factor
Comments	factors describe needs (phil) or relationships (biont) or tolerances (tolerant). azido: acid/chromo: colours/copro: faeces/halo: salt/helio: light/hygro: humidity/hylo: wood/kreno: springs/myceto: fungus/myrmec: ants/necro: carcass/osmo: smell/petro: bolder/pholeo: darkness/psammo: sand/rheo: currents/sapro: decomposition/silico: pebbles/thermo: temperature/tixo: touch/troglo: cave/tyro: bogs_swamps/xero: dryness
example	mycetophil
relationSource	takenfrom
source	Koch (1989) Die Käfer Mitteleuropas - Ökologie. Goecke & Evers Verlag. ISBN 3-87263-037-7

Micro_habitat

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	Micro_habitat
traitName	Micro_habitat
trait_class	environment
broaderTerm	https://ecologicaltraitdata.github.io/TraitDataList/#Ecological_niche
valueType	factor
traitUnit	NA
maxAllowedValue	NA
minAllowedValue	NA
factorLevels	living_tree/litter/faeces/decaying_trunk/tree_hole/ant_hill/mushroom/mineral_surface/roots/corpse_carcass/detritus/under_rocks/moss/sediment
traitDescription	species distribution along an environmental factor describing the microhabitat dimension of the ecological niche
Comments	microhabitat is defined as the smallest part of the environment that substitutes as habitat
example	living_tree
relationSource	equals
source	http://t-sita.cesab.org/BETSI_vizInfo.jsp?trait=Microhabitat

Humidity

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	Humidity
traitName	Humidity
trait_class	environment
broaderTerm	https://ecologicaltraitdata.github.io/TraitDataList/#Ecological_niche
valueType	factor
traitUnit	NA
maxAllowedValue	NA
minAllowedValue	NA
factorLevels	dry/humid/wet
traitDescription	humidity conditions in which individuals occur
Comments	Humidity can be measured in different strata e.g. in the soil or in the air
example	dry
relationSource	broadens
source	http://t-sita.cesab.org/BETSI_vizInfo.jsp?trait=Soil_humidity

Humidity_soil

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	Humidity_soil
traitName	Humidity_soil
trait_class	environment
broaderTerm	https://ecologicaltraitdata.github.io/TraitDataList/#Humidity
valueType	numeric
traitUnit	g/kg
maxAllowedValue	1000
minAllowedValue	0
factorLevels	NA
traitDescription	average value of mass water content in soil at which individuals were observed
Comments
example	10
relationSource	refines
source	http://t-sita.cesab.org/BETSI_vizInfo.jsp?trait=Soil_humidity

Humidity_soil_max

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	Humidity_soil_max
traitName	Humidity_soil_max
trait_class	environment
broaderTerm	https://ecologicaltraitdata.github.io/TraitDataList/#Humidity_soil
valueType	numeric
traitUnit	g/kg
maxAllowedValue	1000
minAllowedValue	0
factorLevels	NA
traitDescription	maximum value of mass water content in soil at which individuals were observed
Comments
example	10
relationSource	inheritsfrom
source	https://ecologicaltraitdata.github.io/TraitDataList/#Humidity_soil

Humidity_soil_min

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	Humidity_soil_min
traitName	Humidity_soil_min
trait_class	environment
broaderTerm	https://ecologicaltraitdata.github.io/TraitDataList/#Humidity_soil
valueType	numeric
traitUnit	g/kg
maxAllowedValue	1000
minAllowedValue	0
factorLevels	NA
traitDescription	minimum value of mass water content in soil at which individuals were observed
Comments
example	0.5
relationSource	inheritsfrom
source	https://ecologicaltraitdata.github.io/TraitDataList/#Humidity_soil

Humidity_air

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	Humidity_air
traitName	Humidity_air
trait_class	environment
broaderTerm	https://ecologicaltraitdata.github.io/TraitDataList/#Humidity
valueType	numeric
traitUnit	%
maxAllowedValue	100
minAllowedValue	0
factorLevels	NA
traitDescription	average relative humidity in the air at which individuals were observed
Comments
example	60
relationSource	inheritsfrom
source	http://t-sita.cesab.org/BETSI_vizInfo.jsp?trait=Soil_humidity

Humidity_air_max

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	Humidity_air_max
traitName	Humidity_air_max
trait_class	environment
broaderTerm	https://ecologicaltraitdata.github.io/TraitDataList/#Humidity_air
valueType	numeric
traitUnit	%
maxAllowedValue	100
minAllowedValue	0
factorLevels	NA
traitDescription	maximum relative humidity in the air at which individuals were observed
Comments
example	60
relationSource	inheritsfrom
source	https://ecologicaltraitdata.github.io/TraitDataList/#Humidity_air

Humidity_air_min

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	Humidity_air_min
traitName	Humidity_air_min
trait_class	environment
broaderTerm	https://ecologicaltraitdata.github.io/TraitDataList/#Humidity_air
valueType	numeric
traitUnit	%
maxAllowedValue	100
minAllowedValue	0
factorLevels	NA
traitDescription	minimum relative humidity in the air at which individuals were observed
Comments
example	5
relationSource	inheritsfrom
source	https://ecologicaltraitdata.github.io/TraitDataList/#Humidity_air

Oxygen_water

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	Oxygen_water
traitName	Oxygen_water
trait_class	environment
broaderTerm	https://ecologicaltraitdata.github.io/TraitDataList/#Ecological_niche
valueType	numeric
traitUnit	mg/L
maxAllowedValue	1000000
minAllowedValue	0
factorLevels	NA
traitDescription	average amount of oxygen in the waterbody at which individuals were observed
Comments
example	10
relationSource
source

Oxygen_water_max

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	Oxygen_water_max
traitName	Oxygen_water_max
trait_class	environment
broaderTerm	https://ecologicaltraitdata.github.io/TraitDataList/#Oxygen_water
valueType	numeric
traitUnit	mg/L
maxAllowedValue	1000000
minAllowedValue	0
factorLevels	NA
traitDescription	maximum amount of oxygen in the waterbody at which individuals were observed
Comments
example	10
relationSource	inheritsfrom
source	https://ecologicaltraitdata.github.io/TraitDataList/#Oxygen_water

Temperature_survival

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	Temperature_survival
traitName	Temperature_survival
trait_class	environment
broaderTerm	https://ecologicaltraitdata.github.io/TraitDataList/#Ecological_niche
valueType	numeric
traitUnit	°C
maxAllowedValue	999
minAllowedValue	-1000
factorLevels	NA
traitDescription	average temperature under which individuals can survive
Comments	Temperature measured in the prefered microhabitat
example	45
relationSource	refines
source	http://t-sita.cesab.org/BETSI_vizInfo.jsp?trait=Soil_temperature

Temperature_survival_max

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	Temperature_survival_max
traitName	Temperature_survival_max
trait_class	environment
broaderTerm	https://ecologicaltraitdata.github.io/TraitDataList/#Temperature_survival
valueType	numeric
traitUnit	°C
maxAllowedValue	999
minAllowedValue	-1000
factorLevels	NA
traitDescription	maximum temperature under which individuals can survive
Comments	Temperature measured in the prefered microhabitat
example	45
relationSource	refines
source	http://t-sita.cesab.org/BETSI_vizInfo.jsp?trait=Soil_temperature

Temperature_survival_min

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	Temperature_survival_min
traitName	Temperature_survival_min
trait_class	environment
broaderTerm	https://ecologicaltraitdata.github.io/TraitDataList/#Temperature_survival
valueType	numeric
traitUnit	°C
maxAllowedValue	999
minAllowedValue	-1000
factorLevels	NA
traitDescription	minimum temperature under which individuals can survive
Comments	Temperature measured in the prefered microhabitat
example	-20
relationSource	refines
source	http://t-sita.cesab.org/BETSI_vizInfo.jsp?trait=Soil_temperature

Temperature_activity

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	Temperature_activity
traitName	Temperature_activity
trait_class	environment
broaderTerm	https://ecologicaltraitdata.github.io/TraitDataList/#Ecological_niche
valueType	numeric
traitUnit	°C
maxAllowedValue	999
minAllowedValue	-1000
factorLevels	NA
traitDescription	average temperature under which individuals are active
Comments	Temperature measured in the prefered microhabitat
example	30
relationSource	refines
source	http://t-sita.cesab.org/BETSI_vizInfo.jsp?trait=Soil_temperature

Temperature_activity_max

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	Temperature_activity_max
traitName	Temperature_activity_max
trait_class	environment
broaderTerm	https://ecologicaltraitdata.github.io/TraitDataList/#Temperature_activity
valueType	numeric
traitUnit	°C
maxAllowedValue	999
minAllowedValue	-1000
factorLevels	NA
traitDescription	maximum temperature under which individuals are active
Comments	Temperature measured in the prefered microhabitat
example	30
relationSource	refines
source	http://t-sita.cesab.org/BETSI_vizInfo.jsp?trait=Soil_temperature

Temperature_activity_min

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	Temperature_activity_min
traitName	Temperature_activity_min
trait_class	environment
broaderTerm	https://ecologicaltraitdata.github.io/TraitDataList/#Temperature_activity
valueType	numeric
traitUnit	°C
maxAllowedValue	999
minAllowedValue	-1000
factorLevels	NA
traitDescription	minimum temperature under which individuals are active
Comments	Temperature measured in the prefered microhabitat
example	10
relationSource	refines
source	http://t-sita.cesab.org/BETSI_vizInfo.jsp?trait=Soil_temperature

Temperature_reproduction

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	Temperature_reproduction
traitName	Temperature_reproduction
trait_class	environment
broaderTerm	https://ecologicaltraitdata.github.io/TraitDataList/#Ecological_niche
valueType	numeric
traitUnit	°C
maxAllowedValue	999
minAllowedValue	-1000
factorLevels	NA
traitDescription	average temperature under which individuals reproduce
Comments	Temperature measured in the prefered microhabitat
example	25
relationSource	refines
source	http://t-sita.cesab.org/BETSI_vizInfo.jsp?trait=Soil_temperature

Temperature_reproduction_max

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	Temperature_reproduction_max
traitName	Temperature_reproduction_max
trait_class	environment
broaderTerm	https://ecologicaltraitdata.github.io/TraitDataList/#Temperature_reproduction
valueType	numeric
traitUnit	°C
maxAllowedValue	999
minAllowedValue	-1000
factorLevels	NA
traitDescription	maximum temperature under which individuals reproduce
Comments	Temperature measured in the prefered microhabitat
example	25
relationSource	refines
source	http://t-sita.cesab.org/BETSI_vizInfo.jsp?trait=Soil_temperature

Temperature_reproduction_min

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	Temperature_reproduction_min
traitName	Temperature_reproduction_min
trait_class	environment
broaderTerm	https://ecologicaltraitdata.github.io/TraitDataList/#Temperature_reproduction
valueType	numeric
traitUnit	°C
maxAllowedValue	999
minAllowedValue	-1000
factorLevels	NA
traitDescription	minimum temperature under which individuals reproduce
Comments	Temperature measured in the prefered microhabitat
example	15
relationSource	refines
source	http://t-sita.cesab.org/BETSI_vizInfo.jsp?trait=Soil_temperature

Temperature_development

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	Temperature_development
traitName	Temperature_development
trait_class	environment
broaderTerm	https://ecologicaltraitdata.github.io/TraitDataList/#Ecological_niche
valueType	numeric
traitUnit	°C
maxAllowedValue	999
minAllowedValue	-1000
factorLevels	NA
traitDescription	average temperature under which juveniles develop into adults
Comments	Temperature measured in the prefered microhabitat
example	30
relationSource	refines
source	http://t-sita.cesab.org/BETSI_vizInfo.jsp?trait=Soil_temperature

Temperature_development_max

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	Temperature_development_max
traitName	Temperature_development_max
trait_class	environment
broaderTerm	https://ecologicaltraitdata.github.io/TraitDataList/#Temperature_development
valueType	numeric
traitUnit	°C
maxAllowedValue	999
minAllowedValue	-1000
factorLevels	NA
traitDescription	maximum temperature under which juveniles develop into adults
Comments	Temperature measured in the prefered microhabitat
example	30
relationSource	refines
source	http://t-sita.cesab.org/BETSI_vizInfo.jsp?trait=Soil_temperature

Temperature_development_min

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	Temperature_development_min
traitName	Temperature_development_min
trait_class	environment
broaderTerm	https://ecologicaltraitdata.github.io/TraitDataList/#Temperature_development
valueType	numeric
traitUnit	°C
maxAllowedValue	999
minAllowedValue	-1000
factorLevels	NA
traitDescription	minimum temperature under which juveniles develop into adults
Comments	Temperature measured in the prefered microhabitat
example	15
relationSource	refines
source	http://t-sita.cesab.org/BETSI_vizInfo.jsp?trait=Soil_temperature

Stratum_preference

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	Stratum_preference
traitName	Stratum_preference
trait_class	environment
broaderTerm	https://ecologicaltraitdata.github.io/TraitDataList/#Habitat_preferences
valueType	factor
traitUnit	NA
maxAllowedValue	NA
minAllowedValue	NA
factorLevels	s/s-(g)/s-(h)/s-(t)/g/(s)-g/g-(h)/g-(t)/h/(s)-h/(g)-h/h-(t)/t/(s)-t/(g)-t/(h)-t
traitDescription	vertical distribution of habitat
Comments	Stratum in which individuals are usually observed. Second prefered habitat is given in brackets. S:soil/g:ground/h:herb-layer/t:shrub & tree_layer
example	ground
relationSource	takenfrom
source	https://www.nature.com/articles/sdata201513

Stratum_phenology

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	Stratum_phenology
traitName	Stratum_phenology
trait_class	environment
broaderTerm	https://ecologicaltraitdata.github.io/TraitDataList/#Stratum_preference
valueType	factor
traitUnit	NA
maxAllowedValue	NA
minAllowedValue	NA
factorLevels	monoroph/allaxoroph/xeroph/monochor/allaxochor/xerochor/homotop/heterotop
traitDescription	individuals occur in the same or different habitats throughout the year
Comments	monoroph: only one stratum/allaxoroph: stratum change/xeroph: several strata at the same time/monochor: at specific locations within one stratum/allaxochor: change of location/xerochor: several locations within one stratum/homotop: complete life-cycle in one habitat/heterotop: life-cycle in different habitats
example	monoroph
relationSource	takenfrom
source	Koch (1989) Die Käfer Mitteleuropas - Ökologie. Goecke & Evers Verlag. ISBN 3-87263-037-7

Habitat_CORINE

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	Habitat_CORINE
traitName	Habitat_CORINE
trait_class	environment
broaderTerm	https://ecologicaltraitdata.github.io/TraitDataList/#Habitat_preferences
valueType	factor
traitUnit	NA
maxAllowedValue	NA
minAllowedValue	NA
factorLevels
traitDescription	habitats where occurrence of individuals is most common
Comments	Land cover classes are based on CORINE land cover classes
example	321-322
relationSource	equals
source	http://t-sita.cesab.org/BETSI_vizInfo.jsp?trait=Habitat

Habitat_forest

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	Habitat_forest
traitName	Habitat_forest
trait_class	environment
broaderTerm	https://ecologicaltraitdata.github.io/TraitDataList/#Habitat_preferences
valueType	logical
traitUnit	NA
maxAllowedValue	NA
minAllowedValue	NA
factorLevels	NA
traitDescription	individuals only occur in forested areas
Comments
example	TRUE
relationSource	equals
source	http://t-sita.cesab.org/BETSI_vizInfo.jsp?trait=Forest

Habitat_grassland

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	Habitat_grassland
traitName	Habitat_grassland
trait_class	environment
broaderTerm	https://ecologicaltraitdata.github.io/TraitDataList/#Habitat_preferences
valueType	logical
traitUnit	NA
maxAllowedValue	NA
minAllowedValue	NA
factorLevels	NA
traitDescription	individuals only occur in grasslands
Comments
example	FALSE
relationSource	equals
source	http://t-sita.cesab.org/BETSI_vizInfo.jsp?trait=Grassland

Nature_Orientation

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	Nature_Orientation
traitName	Nature_Orientation
trait_class	environment
broaderTerm	https://ecologicaltraitdata.github.io/TraitDataList/#Habitat_preferences
valueType	factor
traitUnit	NA
maxAllowedValue	NA
minAllowedValue	NA
factorLevels	climax/semi-natural/disturbed/artificial
traitDescription	occurence of individuals is restricted to habitats with a specific state of naturalness
Comments	naturalness ranges from climax (most natural) to artificial
example	semi-natural
relationSource
source

Climate_zones_Köppen

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	Climate_zones_Köppen
traitName	Climate_zones_Köppen
trait_class	environment
broaderTerm	https://ecologicaltraitdata.github.io/TraitDataList/#Habitat_preferences
valueType	factor
traitUnit	NA
maxAllowedValue	NA
minAllowedValue	NA
factorLevels	Af/Am/As/Aw/BWh/BWk/BSh/BSk/Csa/Csb/Csc/Cwa/Cwb/Cwc/Cfa/Cfb/Cfc/Dsa/Dsb/Dsc/Dsd/Dwa/Dwb/Dwc/Dwd/Dfa/Dfb/Dfc/Dfd/ET/EF
traitDescription	climate zone(s) in which individuals are observed
Comments	Climate zone definition follows Köppen with five main groups: A (tropical)/ B (dry) / C (temperate) / D (continental) / E (polar)
example	Cfb-Csa
relationSource	takenfrom
source	https://en.wikipedia.org/wiki/K%C3%B6ppen_climate_classification

Relict_species

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	Relict_species
traitName	Relict_species
trait_class	environment
broaderTerm	https://ecologicaltraitdata.github.io/TraitDataList/#Habitat_specialization
valueType	logical
traitUnit	NA
maxAllowedValue	NA
minAllowedValue	NA
factorLevels	NA
traitDescription	species is dependent on pre-anthropogenic conditions (e.g. “Urwald-Reliktarten”)
Comments
example	TRUE
relationSource
source

Oxygen_water_min

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	Oxygen_water_min
traitName	Oxygen_water_min
trait_class	environment
broaderTerm	https://ecologicaltraitdata.github.io/TraitDataList/#Oxygen_water
valueType	numeric
traitUnit	mg/L
maxAllowedValue	1000000
minAllowedValue	0
factorLevels	NA
traitDescription	minimum amount of oxygen in the waterbody at which individuals were observed
Comments
example	5
relationSource	inheritsfrom
source	https://ecologicaltraitdata.github.io/TraitDataList/#Oxygen_water

Deadwood_dependent

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	Deadwood_dependent
traitName	Deadwood_dependent
trait_class	environment
broaderTerm	https://ecologicaltraitdata.github.io/TraitDataList/#Habitat_specialization
valueType	logical
traitUnit	NA
maxAllowedValue	NA
minAllowedValue	NA
factorLevels	NA
traitDescription	species is obligat dependent on deadwood
Comments	Dependence needs to be critical for survival or reproduction
example	TRUE
relationSource
source

Traits relating to feeding

Stable_Isotopes

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	Stable_Isotopes
traitName	Stable_Isotopes
trait_class	feeding
broaderTerm	https://ecologicaltraitdata.github.io/TraitDataList/#Stoichiometry
valueType	numeric
traitUnit	delta per mill
maxAllowedValue	NA
minAllowedValue	NA
factorLevels	NA
traitDescription
Comments
example
relationSource
source

Feeding_guild

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	Feeding_guild
traitName	Feeding_guild
trait_class	feeding
broaderTerm	https://ecologicaltraitdata.github.io/TraitDataList/#feeding
valueType	factor
traitUnit	NA
maxAllowedValue	NA
minAllowedValue	NA
factorLevels	h/c/f/d/o
traitDescription	feeding guild of the species
Comments	h: herbivor/ c: carnivor/ f: fungivores/ d: detritivor/ o: omnivores
example	c
relationSource	takenfrom
source	https://www.nature.com/articles/sdata201513

Feeding_guild_adults

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	Feeding_guild_adults
traitName	Feeding_guild_adults
trait_class	feeding
broaderTerm	https://ecologicaltraitdata.github.io/TraitDataList/#Feeding_guild
valueType	factor
traitUnit	NA
maxAllowedValue	NA
minAllowedValue	NA
factorLevels	h/c/f/d/o
traitDescription	feeding guild of the species in its adult stage
Comments	h: herbivor/ c: carnivor/ f: fungivores/ d: detritivor/ o: omnivores
example	c
relationSource	takenfrom
source	https://www.nature.com/articles/sdata201513

Feeding_guild_juveniles

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	Feeding_guild_juveniles
traitName	Feeding_guild_juveniles
trait_class	feeding
broaderTerm	https://ecologicaltraitdata.github.io/TraitDataList/#Feeding_guild
valueType	factor
traitUnit	NA
maxAllowedValue	NA
minAllowedValue	NA
factorLevels	h/c/f/d/o
traitDescription	feeding guild of the species in its juvenile stage
Comments	h: herbivor/ c: carnivor/ f: fungivores/ d: detritivor/ o: omnivores
example	c
relationSource	takenfrom
source	https://www.nature.com/articles/sdata201513

Feeding_mode

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	Feeding_mode
traitName	Feeding_mode
trait_class	feeding
broaderTerm	https://ecologicaltraitdata.github.io/TraitDataList/#Feeding_guild
valueType	factor
traitUnit	NA
maxAllowedValue	NA
minAllowedValue	NA
factorLevels	e/c/s
traitDescription	mode of aquiring the nutrients
Comments	e: extraintestinal/ c: chewing/ s:sucking
example	e
relationSource	takenfrom
source	https://www.nature.com/articles/sdata201513

Feeding_mode_juveniles

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	Feeding_mode_juveniles
traitName	Feeding_mode_juveniles
trait_class	feeding
broaderTerm	https://ecologicaltraitdata.github.io/TraitDataList/#Feeding_guild
valueType	factor
traitUnit	NA
maxAllowedValue	NA
minAllowedValue	NA
factorLevels	e/c/s
traitDescription	mode of aquiring the nutrients during larval stage
Comments	e: extraintestinal/ c: chewing/ s:sucking
example	e
relationSource	takenfrom
source	https://www.nature.com/articles/sdata201513

Feeding_specialization

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	Feeding_specialization
traitName	Feeding_specialization
trait_class	feeding
broaderTerm	https://ecologicaltraitdata.github.io/TraitDataList/#Feeding_guild
valueType	factor
traitUnit	NA
maxAllowedValue	NA
minAllowedValue	NA
factorLevels	m/o/p
traitDescription	width of host specialization
Comments	m: monophag (on one species or genus of host)/ o: oligophag (on one family or subfamily of host)/ p: poliphag (on more than one family of host)
example	p
relationSource	takenfrom
source	https://www.nature.com/articles/sdata201513

Host

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	Host
traitName	Host
trait_class	feeding
broaderTerm	https://ecologicaltraitdata.github.io/TraitDataList/#Feeding_specialization
valueType	character
traitUnit	NA
maxAllowedValue	NA
minAllowedValue	NA
factorLevels	NA
traitDescription	scientific name or group of hosts or prey
Comments
example	Quercus robur
relationSource
source

Host_number

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	Host_number
traitName	Host_number
trait_class	feeding
broaderTerm	https://ecologicaltraitdata.github.io/TraitDataList/#Host
valueType	integer
traitUnit	count
maxAllowedValue	9999
minAllowedValue	1
factorLevels	NA
traitDescription	number of different known host or prey species
Comments
example	5
relationSource
source

Endoghagous_lifestile

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	Endoghagous_lifestile
traitName	Endoghagous_lifestile
trait_class	feeding
broaderTerm	https://ecologicaltraitdata.github.io/TraitDataList/#Feeding_specialization
valueType	factor
traitUnit	NA
maxAllowedValue	NA
minAllowedValue	NA
factorLevels	gl/gr/gs/ml/mr/ms/mro
traitDescription	details on endophagously living larvae
Comments	gl: gall-inducer leaves / g: gall-inducer reproductive organs / gs: gall-inducer stems / ml: miner leaves / mr: miner reproductive organs / ms: miner stem / mro: miner roots
example	ml
relationSource	takenfrom
source	https://www.nature.com/articles/sdata201513

Feeding_specialization_juveniles

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	Feeding_specialization_juveniles
traitName	Feeding_specialization_juveniles
trait_class	feeding
broaderTerm	https://ecologicaltraitdata.github.io/TraitDataList/#Feeding_guild
valueType	factor
traitUnit	NA
maxAllowedValue	NA
minAllowedValue	NA
factorLevels	m/o/p
traitDescription	width of host specialization in juveniles
Comments	m: monophag (on one species or genus of host)/ o: oligophag (on one family or subfamily of host)/ p: poliphag (on more than one family of host)
example	p
relationSource	takenfrom
source	https://www.nature.com/articles/sdata201513

Feeding_tissue

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	Feeding_tissue
traitName	Feeding_tissue
trait_class	feeding
broaderTerm	https://ecologicaltraitdata.github.io/TraitDataList/#Feeding_guild
valueType	factor
traitUnit	NA
maxAllowedValue	NA
minAllowedValue	NA
factorLevels	m/p/r/x/se/bl/bo/f/ms/w
traitDescription	tissue from which nutrients are aquired
Comments	m: mesophyll-cells / p: phloem / r: reproductive organs / x: xylem / se: ripe seeds / bl: blood / bo: bones / f: fatty tissue / ms: muscular tissue / w: whole organism
example	bl
relationSource	refines
source	https://www.nature.com/articles/sdata201513

Feeding_plant_part

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	Feeding_plant_part
traitName	Feeding_plant_part
trait_class	feeding
broaderTerm	https://ecologicaltraitdata.github.io/TraitDataList/#Feeding_tissue
valueType	factor
traitUnit	NA
maxAllowedValue	NA
minAllowedValue	NA
factorLevels	l/r/ro/s/se
traitDescription	plant parts on which herbivorous species feed
Comments	l: leaves / r: reproductive organs / ro: roots / s: stem / se: ripe seeds
example	l
relationSource	takenfrom
source	https://www.nature.com/articles/sdata201513

Feeding_plant_part_juveniles

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	Feeding_plant_part_juveniles
traitName	Feeding_plant_part_juveniles
trait_class	feeding
broaderTerm	https://ecologicaltraitdata.github.io/TraitDataList/#Feeding_tissue
valueType	factor
traitUnit	NA
maxAllowedValue	NA
minAllowedValue	NA
factorLevels	l/r/ro/s/se
traitDescription	plant parts on which herbivorous species feed during juvenile stage
Comments	l: leaves / r: reproductive organs / ro: roots / s: stem / se: ripe seeds
example	l
relationSource	takenfrom
source	https://www.nature.com/articles/sdata201513

Feeding_tissue_juveniles

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	Feeding_tissue_juveniles
traitName	Feeding_tissue_juveniles
trait_class	feeding
broaderTerm	https://ecologicaltraitdata.github.io/TraitDataList/#Feeding_guild
valueType	factor
traitUnit	NA
maxAllowedValue	NA
minAllowedValue	NA
factorLevels	m/p/r/x/se/bl/bo/f/ms/w
traitDescription	tissue from which nutrients are aquired during the juvenil stage
Comments	m: mesophyll-cells / p: phloem / r: reproductive organs / x: xylem / se: ripe seeds / bl: blood / bo: bones / f: fatty tissue / ms: muscular tissue / w: whole organism
example	bl
relationSource	refines
source	https://www.nature.com/articles/sdata201513

Feeding_behaviour

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	Feeding_behaviour
traitName	Feeding_behaviour
trait_class	feeding
broaderTerm	https://ecologicaltraitdata.github.io/TraitDataList/#Feeding_guild
valueType	factor
traitUnit	NA
maxAllowedValue	NA
minAllowedValue	NA
factorLevels	trap_building/active_hunting/stalking/ambushing/foraging/scavening/grazing/run_and_kill/sit_and_wait
traitDescription	strategy to aquire food items
Comments
example	stalking
relationSource	refines
source	http://t-sita.cesab.org/BETSI_vizInfo.jsp?trait=Hunting_strategy

Trap_strategy

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	Trap_strategy
traitName	Trap_strategy
trait_class	feeding
broaderTerm	https://ecologicaltraitdata.github.io/TraitDataList/#Feeding_behaviour
valueType	factor
traitUnit	NA
maxAllowedValue	NA
minAllowedValue	NA
factorLevels	web/orb/funnel
traitDescription	type of trap constructed to catch prey
Comments
example	web
relationSource	refines
source	http://t-sita.cesab.org/BETSI_vizInfo.jsp?trait=Trap_strategy

Trophic_level

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	Trophic_level
traitName	Trophic_level
trait_class	feeding
broaderTerm	https://ecologicaltraitdata.github.io/TraitDataList/#feeding
valueType	factor
traitUnit	NA
maxAllowedValue	NA
minAllowedValue	NA
factorLevels	producer/consumer/consumer_higher_level/destruent
traitDescription	general trophic level
Comments
example	consumer
relationSource
source

Trophic_level_num

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	Trophic_level_num
traitName	Trophic_level_num
trait_class	feeding
broaderTerm	https://ecologicaltraitdata.github.io/TraitDataList/#Trophic_level
valueType	numeric
traitUnit	level
maxAllowedValue	999
minAllowedValue	-1
factorLevels	NA
traitDescription	trophic level within the foodweb
Comments	levels are counted as -1: destruent /0: producers /1: first level consumer / 2: second level consumer
example	2
relationSource
source

Traits describing the life-history

Sexuality

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	Sexuality
traitName	Sexuality
trait_class	life-history
broaderTerm	https://ecologicaltraitdata.github.io/TraitDataList/#life-history
valueType	factor
traitUnit	NA
maxAllowedValue	NA
minAllowedValue	NA
factorLevels	clonal/parthenogenetic/sexual/asexual
traitDescription	strategy for transfer of genes
Comments
example	parthenogentic
relationSource	refines
source	http://t-sita.cesab.org/BETSI_vizInfo.jsp?trait=Reproduction_type

Asexual_strategy

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	Asexual_strategy
traitName	Asexual_strategy
trait_class	life-history
broaderTerm	https://ecologicaltraitdata.github.io/TraitDataList/#Sexuality
valueType	factor
traitUnit	NA
maxAllowedValue	NA
minAllowedValue	NA
factorLevels	amphytokus/arrhenotokus/deuterotokus/thelyotokus
traitDescription	mode of asexual reproduction
Comments	if genes are only transfered from one parent to the offspring
example	amphytokus
relationSource	equals
source	http://t-sita.cesab.org/BETSI_vizInfo.jsp?trait=Asexual_reproduction

Reproduction_mode

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	Reproduction_mode
traitName	Reproduction_mode
trait_class	life-history
broaderTerm	https://ecologicaltraitdata.github.io/TraitDataList/#life-history
valueType	factor
traitUnit	NA
maxAllowedValue	NA
minAllowedValue	NA
factorLevels	ovipar/vivipar/ovo-vivipar/ovulipar/histotrophic_vivipar/hemotrophic_vivipar
traitDescription	mode of reprodruction
Comments	describes the state in which offspring are produced by the adults
example	ovipar
relationSource	refines
source	http://t-sita.cesab.org/BETSI_vizInfo.jsp?trait=Carrying_eggs%20strategy

Reproduction_strategy

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	Reproduction_strategy
traitName	Reproduction_strategy
trait_class	life-history
broaderTerm	https://ecologicaltraitdata.github.io/TraitDataList/#Reproduction_mode
valueType	factor
traitUnit	NA
maxAllowedValue	NA
minAllowedValue	NA
factorLevels	iteroparous/semelparous
traitDescription	strategy for number of reproduction events before death
Comments	iteroparous: multiple events during life-time / semelparous: one event during life-span
example	iteroparous
relationSource	refines
source	http://t-sita.cesab.org/BETSI_vizInfo.jsp?trait=Reproduction_behaviour

Clutch_size

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	Clutch_size
traitName	Clutch_size
trait_class	life-history
broaderTerm	https://ecologicaltraitdata.github.io/TraitDataList/#Reproduction_mode
valueType	numeric
traitUnit	count
maxAllowedValue	9999
minAllowedValue	1
factorLevels	NA
traitDescription	number of eggs in a clutch
Comments	A clutch is a group of eggs which are released or laid within a defined time frame or cared for simultaneously (e.g. within one nest)
example	5
relationSource	refines
source	http://t-sita.cesab.org/BETSI_vizInfo.jsp?trait=Clutch_length

Clutch_size_max

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	Clutch_size_max
traitName	Clutch_size_max
trait_class	life-history
broaderTerm	https://ecologicaltraitdata.github.io/TraitDataList/#Clutch_size
valueType	numeric
traitUnit	count
maxAllowedValue	9999
minAllowedValue	1
factorLevels	NA
traitDescription	maximum number of eggs in a clutch
Comments	A clutch is a group of eggs which are released or laid within a defined time frame or cared for simultaneously (e.g. within one nest)
example	5
relationSource	refines
source	https://ecologicaltraitdata.github.io/TraitDataList/#Clutch_size

Clutch_size_min

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	Clutch_size_min
traitName	Clutch_size_min
trait_class	life-history
broaderTerm	https://ecologicaltraitdata.github.io/TraitDataList/#Clutch_size
valueType	numeric
traitUnit	count
maxAllowedValue	9999
minAllowedValue	1
factorLevels	NA
traitDescription	minimum number of eggs in a clutch
Comments	A clutch is a group of eggs which are released or laid within a defined time frame or cared for simultaneously (e.g. within one nest)
example	5
relationSource	refines
source	https://ecologicaltraitdata.github.io/TraitDataList/#Clutch_size

Nr_eggs_lifetime

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	Nr_eggs_lifetime
traitName	Nr_eggs_lifetime
trait_class	life-history
broaderTerm	https://ecologicaltraitdata.github.io/TraitDataList/#Reproduction_mode
valueType	numeric
traitUnit	count
maxAllowedValue	9999
minAllowedValue	1
factorLevels	NA
traitDescription	number of eggs produced by one individual over their life-span
Comments
example	500
relationSource
source

Nr_eggs_lifetime_max

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	Nr_eggs_lifetime_max
traitName	Nr_eggs_lifetime_max
trait_class	life-history
broaderTerm	https://ecologicaltraitdata.github.io/TraitDataList/#Nr_eggs_lifetime
valueType	numeric
traitUnit	count
maxAllowedValue	9999
minAllowedValue	1
factorLevels	NA
traitDescription	maximum number of eggs produced by one individual over their life-span
Comments
example	500
relationSource	inheritsfrom
source	https://ecologicaltraitdata.github.io/TraitDataList/#Nr_eggs_lifetime

Nr_eggs_lifetime_min

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	Nr_eggs_lifetime_min
traitName	Nr_eggs_lifetime_min
trait_class	life-history
broaderTerm	https://ecologicaltraitdata.github.io/TraitDataList/#Nr_eggs_lifetime
valueType	numeric
traitUnit	count
maxAllowedValue	9999
minAllowedValue	1
factorLevels	NA
traitDescription	minimum number of eggs produced by one individual over their life-span
Comments
example	500
relationSource	inheritsfrom
source	https://ecologicaltraitdata.github.io/TraitDataList/#Nr_eggs_lifetime

Egg_size

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	Egg_size
traitName	Egg_size
trait_class	life-history
broaderTerm	https://ecologicaltraitdata.github.io/TraitDataList/#Reproduction_mode
valueType	factor
traitUnit	NA
maxAllowedValue	NA
minAllowedValue	NA
factorLevels	<1mm/<1cm/<10cm/<100m
traitDescription	size class of indivdiual eggs
Comments
example	<1cm
relationSource	refines
source	http://t-sita.cesab.org/BETSI_vizInfo.jsp?trait=Clutch_length

Egg_length

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	Egg_length
traitName	Egg_length
trait_class	life-history
broaderTerm	https://ecologicaltraitdata.github.io/TraitDataList/#Egg_size
valueType	numeric
traitUnit	cm
maxAllowedValue	999
minAllowedValue	0
factorLevels	NA
traitDescription	length of individual eggs
Comments	measured along the longest axis in oval shapes
example	0.2
relationSource	refines
source	https://ecologicaltraitdata.github.io/TraitDataList/#Egg_size

Egg_length_max

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	Egg_length_max
traitName	Egg_length_max
trait_class	life-history
broaderTerm	https://ecologicaltraitdata.github.io/TraitDataList/#Egg_length
valueType	numeric
traitUnit	cm
maxAllowedValue	999
minAllowedValue	0
factorLevels	NA
traitDescription	maximum observed length of individual eggs
Comments	measured along the longest axis in oval shapes
example	1.5
relationSource	inheritsfrom
source	https://ecologicaltraitdata.github.io/TraitDataList/#Egg_length

Egg_length_min

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	Egg_length_min
traitName	Egg_length_min
trait_class	life-history
broaderTerm	https://ecologicaltraitdata.github.io/TraitDataList/#Egg_length
valueType	numeric
traitUnit	cm
maxAllowedValue	999
minAllowedValue	0
factorLevels	NA
traitDescription	minimum observed length of individual eggs
Comments	measured along the longest axis in oval shapes
example	0.1
relationSource	inheritsfrom
source	https://ecologicaltraitdata.github.io/TraitDataList/#Egg_length

Egg_width

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	Egg_width
traitName	Egg_width
trait_class	life-history
broaderTerm	https://ecologicaltraitdata.github.io/TraitDataList/#Egg_size
valueType	numeric
traitUnit	cm
maxAllowedValue	999
minAllowedValue	0
factorLevels	NA
traitDescription	width of individual eggs
Comments	measured along the shorter axis in oval shapes
example	0.2
relationSource	refines
source	https://ecologicaltraitdata.github.io/TraitDataList/#Egg_size

Egg_width_max

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	Egg_width_max
traitName	Egg_width_max
trait_class	life-history
broaderTerm	https://ecologicaltraitdata.github.io/TraitDataList/#Egg_width
valueType	numeric
traitUnit	cm
maxAllowedValue	999
minAllowedValue	0
factorLevels	NA
traitDescription	maximum observed width of individual eggs
Comments	measured along the shorter axis in oval shapes
example	1.5
relationSource	inheritsfrom
source	https://ecologicaltraitdata.github.io/TraitDataList/#Egg_width

Egg_width_min

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	Egg_width_min
traitName	Egg_width_min
trait_class	life-history
broaderTerm	https://ecologicaltraitdata.github.io/TraitDataList/#Egg_width
valueType	numeric
traitUnit	cm
maxAllowedValue	999
minAllowedValue	0
factorLevels	NA
traitDescription	minimum observed width of individual eggs
Comments	measured along the shorter axis in oval shapes
example	0.1
relationSource	inheritsfrom
source	https://ecologicaltraitdata.github.io/TraitDataList/#Egg_width

Egg_diameter

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	Egg_diameter
traitName	Egg_diameter
trait_class	life-history
broaderTerm	https://ecologicaltraitdata.github.io/TraitDataList/#Egg_size
valueType	numeric
traitUnit	cm
maxAllowedValue	999
minAllowedValue	0
factorLevels	NA
traitDescription	diameter of individual eggs
Comments
example	0.2
relationSource	refines
source	https://ecologicaltraitdata.github.io/TraitDataList/#Egg_size

Egg_diameter_max

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	Egg_diameter_max
traitName	Egg_diameter_max
trait_class	life-history
broaderTerm	https://ecologicaltraitdata.github.io/TraitDataList/#Egg_diameter
valueType	numeric
traitUnit	cm
maxAllowedValue	999
minAllowedValue	0
factorLevels	NA
traitDescription	maximum observed diameter of individual eggs
Comments
example	1.5
relationSource	inheritsfrom
source	https://ecologicaltraitdata.github.io/TraitDataList/#Egg_diameter

Egg_diameter_min

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	Egg_diameter_min
traitName	Egg_diameter_min
trait_class	life-history
broaderTerm	https://ecologicaltraitdata.github.io/TraitDataList/#Egg_diameter
valueType	numeric
traitUnit	cm
maxAllowedValue	999
minAllowedValue	0
factorLevels	NA
traitDescription	minimum observed diameter of individual eggs
Comments
example	0.1
relationSource	inheritsfrom
source	https://ecologicaltraitdata.github.io/TraitDataList/#Egg_diameter

Egg_volume

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	Egg_volume
traitName	Egg_volume
trait_class	life-history
broaderTerm	https://ecologicaltraitdata.github.io/TraitDataList/#Egg_size
valueType	numeric
traitUnit	cm³
maxAllowedValue	9999
minAllowedValue	0
factorLevels	NA
traitDescription	volume of individual eggs
Comments
example	0.125
relationSource	refines
source	https://ecologicaltraitdata.github.io/TraitDataList/#Egg_size

Egg_volume_max

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	Egg_volume_max
traitName	Egg_volume_max
trait_class	life-history
broaderTerm	https://ecologicaltraitdata.github.io/TraitDataList/#Egg_volume
valueType	numeric
traitUnit	cm³
maxAllowedValue	9999
minAllowedValue	0
factorLevels	NA
traitDescription	maximum observed volume of individual eggs
Comments
example	0.75
relationSource	inheritsfrom
source	https://ecologicaltraitdata.github.io/TraitDataList/#Egg_volume

Egg_volume_min

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	Egg_volume_min
traitName	Egg_volume_min
trait_class	life-history
broaderTerm	https://ecologicaltraitdata.github.io/TraitDataList/#Egg_volume
valueType	numeric
traitUnit	cm³
maxAllowedValue	9999
minAllowedValue	0
factorLevels	NA
traitDescription	minimum observed volume of individual eggs
Comments
example	0.1
relationSource	inheritsfrom
source	https://ecologicaltraitdata.github.io/TraitDataList/#Egg_volume

Egg_weight

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	Egg_weight
traitName	Egg_weight
trait_class	life-history
broaderTerm	https://ecologicaltraitdata.github.io/TraitDataList/#Egg_size
valueType	numeric
traitUnit	g
maxAllowedValue	999
minAllowedValue	0
factorLevels	NA
traitDescription	weight of individual eggs
Comments
example	2.5
relationSource	refines
source	https://ecologicaltraitdata.github.io/TraitDataList/#Egg_size

Egg_weight_max

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	Egg_weight_max
traitName	Egg_weight_max
trait_class	life-history
broaderTerm	https://ecologicaltraitdata.github.io/TraitDataList/#Egg_weight
valueType	numeric
traitUnit	g
maxAllowedValue	999
minAllowedValue	0
factorLevels	NA
traitDescription	maximum observed weight of individual eggs
Comments
example	2.5
relationSource	inheritsfrom
source	https://ecologicaltraitdata.github.io/TraitDataList/#Egg_weight

Egg_weight_min

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	Egg_weight_min
traitName	Egg_weight_min
trait_class	life-history
broaderTerm	https://ecologicaltraitdata.github.io/TraitDataList/#Egg_weight
valueType	numeric
traitUnit	g
maxAllowedValue	999
minAllowedValue	0
factorLevels	NA
traitDescription	minimum observed weight of individual eggs
Comments
example	2.5
relationSource	inheritsfrom
source	https://ecologicaltraitdata.github.io/TraitDataList/#Egg_weight

Egg_biomass

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	Egg_biomass
traitName	Egg_biomass
trait_class	life-history
broaderTerm	https://ecologicaltraitdata.github.io/TraitDataList/#Egg_size
valueType	numeric
traitUnit	g
maxAllowedValue	999
minAllowedValue	0
factorLevels	NA
traitDescription	estimated biomass of individual eggs
Comments	Estimates can be based on volume or size measurements
example	1.5
relationSource	refines
source	https://ecologicaltraitdata.github.io/TraitDataList/#Egg_size

Egg_development_time

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	Egg_development_time
traitName	Egg_development_time
trait_class	life-history
broaderTerm	https://ecologicaltraitdata.github.io/TraitDataList/#Reproduction_mode
valueType	numeric
traitUnit	day
maxAllowedValue	999
minAllowedValue	0
factorLevels	NA
traitDescription	time between fertilization and hatching
Comments
example	5
relationSource	refines
source	http://t-sita.cesab.org/BETSI_vizInfo.jsp?trait=Development

Egg_development_time_max

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	Egg_development_time_max
traitName	Egg_development_time_max
trait_class	life-history
broaderTerm	https://ecologicaltraitdata.github.io/TraitDataList/#Egg_development_time
valueType	numeric
traitUnit	day
maxAllowedValue	999
minAllowedValue	0
factorLevels	NA
traitDescription	maximum observed time between fertilization and hatching
Comments
example	15
relationSource	inheritsfrom
source	https://ecologicaltraitdata.github.io/TraitDataList/#Egg_development_time

Egg_development_time_min

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	Egg_development_time_min
traitName	Egg_development_time_min
trait_class	life-history
broaderTerm	https://ecologicaltraitdata.github.io/TraitDataList/#Egg_development_time
valueType	numeric
traitUnit	day
maxAllowedValue	999
minAllowedValue	0
factorLevels	NA
traitDescription	minimum observed time between fertilization and hatching
Comments
example	3
relationSource	inheritsfrom
source	https://ecologicaltraitdata.github.io/TraitDataList/#Egg_development_time

Developmental_stages

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	Developmental_stages
traitName	Developmental_stages
trait_class	life-history
broaderTerm	https://ecologicaltraitdata.github.io/TraitDataList/#Reproduction_mode
valueType	factor
traitUnit	NA
maxAllowedValue	NA
minAllowedValue	NA
factorLevels	holometabol/hemimetabol/ametabol
traitDescription	mode of metamorphosis
Comments	describes development of juveniles between hatching and maturity
example	holometabol
relationSource	equals
source	http://t-sita.cesab.org/BETSI_vizInfo.jsp?trait=Stadial_number

Developmental_stages_number

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	Developmental_stages_number
traitName	Developmental_stages_number
trait_class	life-history
broaderTerm	https://ecologicaltraitdata.github.io/TraitDataList/#Developmental_stages
valueType	numeric
traitUnit	count
maxAllowedValue	999
minAllowedValue	0
factorLevels	NA
traitDescription	number of distinct metabolic stages during maturation
Comments
example	7
relationSource	refines
source	https://ecologicaltraitdata.github.io/TraitDataList/#Developmental_stages

Nr_larval_stages

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	Nr_larval_stages
traitName	Nr_larval_stages
trait_class	life-history
broaderTerm	https://ecologicaltraitdata.github.io/TraitDataList/#Developmental_stages_number
valueType	numeric
traitUnit	count
maxAllowedValue	999
minAllowedValue	0
factorLevels	NA
traitDescription	number of larval stages
Comments
example	3
relationSource	refines
source	http://t-sita.cesab.org/BETSI_vizInfo.jsp?trait=Stadial_number

Developmental_stages_time

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	Developmental_stages_time
traitName	Developmental_stages_time
trait_class	life-history
broaderTerm	https://ecologicaltraitdata.github.io/TraitDataList/#Developmental_stages
valueType	numeric
traitUnit	day
maxAllowedValue	999
minAllowedValue	0
factorLevels	NA
traitDescription	time between hatching and maturation
Comments
example	15
relationSource	equals
source	http://t-sita.cesab.org/BETSI_vizInfo.jsp?trait=Juvenile_stadia_duration

Developmental_stages_time_max

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	Developmental_stages_time_max
traitName	Developmental_stages_time_max
trait_class	life-history
broaderTerm	https://ecologicaltraitdata.github.io/TraitDataList/#Developmental_stages_time
valueType	numeric
traitUnit	day
maxAllowedValue	999
minAllowedValue	0
factorLevels	NA
traitDescription	maximum observed time between hatching and maturation
Comments
example	15
relationSource	inheritsfrom
source	https://ecologicaltraitdata.github.io/TraitDataList/#Developmental_stages_time

Developmental_stages_time_min

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	Developmental_stages_time_min
traitName	Developmental_stages_time_min
trait_class	life-history
broaderTerm	https://ecologicaltraitdata.github.io/TraitDataList/#Developmental_stages_time
valueType	numeric
traitUnit	day
maxAllowedValue	999
minAllowedValue	0
factorLevels	NA
traitDescription	minimum observed time between hatching and maturation
Comments
example	15
relationSource	inheritsfrom
source	https://ecologicaltraitdata.github.io/TraitDataList/#Developmental_stages_time

Fecundity

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	Fecundity
traitName	Fecundity
trait_class	life-history
broaderTerm	https://ecologicaltraitdata.github.io/TraitDataList/#Reproduction_mode
valueType	factor
traitUnit	NA
maxAllowedValue	NA
minAllowedValue	NA
factorLevels	low/medium/high
traitDescription	level of fecundity
Comments
example	medium
relationSource
source

Generation_number_year

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	Generation_number_year
traitName	Generation_number_year
trait_class	life-history
broaderTerm	https://ecologicaltraitdata.github.io/TraitDataList/#Fecundity
valueType	numeric
traitUnit	count
maxAllowedValue	999
minAllowedValue	0
factorLevels	NA
traitDescription	number of generations which are produced within one year
Comments
example	2
relationSource	refines
source	http://t-sita.cesab.org/BETSI_vizInfo.jsp?trait=Voltinism

Generation_number_year_max

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	Generation_number_year_max
traitName	Generation_number_year_max
trait_class	life-history
broaderTerm	https://ecologicaltraitdata.github.io/TraitDataList/#Generation_number_year
valueType	numeric
traitUnit	count
maxAllowedValue	999
minAllowedValue	0
factorLevels	NA
traitDescription	maximum observed number of generations which are produced within one year
Comments
example	4
relationSource	inheritsfrom
source	https://ecologicaltraitdata.github.io/TraitDataList/#Generation_number_year

Generation_number_year_min

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	Generation_number_year_min
traitName	Generation_number_year_min
trait_class	life-history
broaderTerm	https://ecologicaltraitdata.github.io/TraitDataList/#Generation_number_year
valueType	numeric
traitUnit	count
maxAllowedValue	999
minAllowedValue	0
factorLevels	NA
traitDescription	minimum observed number of generations which are produced within one year
Comments
example	1
relationSource	inheritsfrom
source	https://ecologicaltraitdata.github.io/TraitDataList/#Generation_number_year

Reproduction_events_year

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	Reproduction_events_year
traitName	Reproduction_events_year
trait_class	life-history
broaderTerm	https://ecologicaltraitdata.github.io/TraitDataList/#Fecundity
valueType	numeric
traitUnit	count
maxAllowedValue	999
minAllowedValue	0
factorLevels	NA
traitDescription	number of reproduction events per individual within one year or season
Comments
example	2
relationSource	equals
source	http://t-sita.cesab.org/BETSI_vizInfo.jsp?trait=Voltinism

Reproduction_events_year_max

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	Reproduction_events_year_max
traitName	Reproduction_events_year_max
trait_class	life-history
broaderTerm	https://ecologicaltraitdata.github.io/TraitDataList/#Reproduction_events_year
valueType	numeric
traitUnit	count
maxAllowedValue	999
minAllowedValue	0
factorLevels	NA
traitDescription	maximum observed number of reproduction events per individual within one year or season
Comments
example	4
relationSource	inheritsfrom
source	https://ecologicaltraitdata.github.io/TraitDataList/#Reproduction_events_year

Reproduction_events_year_min

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	Reproduction_events_year_min
traitName	Reproduction_events_year_min
trait_class	life-history
broaderTerm	https://ecologicaltraitdata.github.io/TraitDataList/#Reproduction_events_year
valueType	numeric
traitUnit	count
maxAllowedValue	999
minAllowedValue	0
factorLevels	NA
traitDescription	minimum observed number of reproduction events per individual within one year or season
Comments
example	1
relationSource	inheritsfrom
source	https://ecologicaltraitdata.github.io/TraitDataList/#Reproduction_events_year

Reproduction_events_life

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	Reproduction_events_life
traitName	Reproduction_events_life
trait_class	life-history
broaderTerm	https://ecologicaltraitdata.github.io/TraitDataList/#Fecundity
valueType	numeric
traitUnit	count
maxAllowedValue	999
minAllowedValue	0
factorLevels	NA
traitDescription	number of reproduction events per individual during ist life-span
Comments
example	2
relationSource	inheritsfrom
source	http://t-sita.cesab.org/BETSI_vizInfo.jsp?trait=Voltinism

Reproduction_events_life_max

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	Reproduction_events_life_max
traitName	Reproduction_events_life_max
trait_class	life-history
broaderTerm	https://ecologicaltraitdata.github.io/TraitDataList/#Reproduction_events_life
valueType	numeric
traitUnit	count
maxAllowedValue	999
minAllowedValue	0
factorLevels	NA
traitDescription	maximum observed number of reproduction events per individual during ist life-span
Comments
example	2
relationSource	inheritsfrom
source	https://ecologicaltraitdata.github.io/TraitDataList/#Reproduction_events_life

Reproduction_events_life_min

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	Reproduction_events_life_min
traitName	Reproduction_events_life_min
trait_class	life-history
broaderTerm	https://ecologicaltraitdata.github.io/TraitDataList/#Reproduction_events_life
valueType	numeric
traitUnit	count
maxAllowedValue	999
minAllowedValue	0
factorLevels	NA
traitDescription	minimum observed number of reproduction events per individual during ist life-span
Comments
example	2
relationSource	inheritsfrom
source	https://ecologicaltraitdata.github.io/TraitDataList/#Reproduction_events_life

Mating_system

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	Mating_system
traitName	Mating_system
trait_class	life-history
broaderTerm	https://ecologicaltraitdata.github.io/TraitDataList/#life-history
valueType	factor
traitUnit	NA
maxAllowedValue	NA
minAllowedValue	NA
factorLevels	polyandry/polygamy/polygynandry/promiscuity
traitDescription	system of mate choice
Comments
example	polyandry
relationSource	refines
source	http://t-sita.cesab.org/BETSI_vizInfo.jsp?trait=Sociability

Life_span_potential

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	Life_span_potential
traitName	Life_span_potential
trait_class	life-history
broaderTerm	https://ecologicaltraitdata.github.io/TraitDataList/#life-history
valueType	numeric
traitUnit	day
maxAllowedValue	9999
minAllowedValue	0
factorLevels	NA
traitDescription	average time between hatching and death due to age
Comments	length of life-span under optimal conditions excluding negative biotic and abiotic interactions
example	90
relationSource	refines
source	http://t-sita.cesab.org/BETSI_vizInfo.jsp?trait=Lifespan

Life_span_potential_max

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	Life_span_potential_max
traitName	Life_span_potential_max
trait_class	life-history
broaderTerm	https://ecologicaltraitdata.github.io/TraitDataList/#Life_span_potential
valueType	numeric
traitUnit	day
maxAllowedValue	9999
minAllowedValue	0
factorLevels	NA
traitDescription	maximum observed time between hatching and death due to age
Comments	length of life-span under optimal conditions excluding negative biotic and abiotic interactions
example	150
relationSource	inheritsfrom
source	https://ecologicaltraitdata.github.io/TraitDataList/#Life_span_potential

Life_span_potential_min

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	Life_span_potential_min
traitName	Life_span_potential_min
trait_class	life-history
broaderTerm	https://ecologicaltraitdata.github.io/TraitDataList/#Life_span_potential
valueType	numeric
traitUnit	day
maxAllowedValue	9999
minAllowedValue	0
factorLevels	NA
traitDescription	minimum observed time between hatching and death due to age
Comments	length of life-span under optimal conditions excluding negative biotic and abiotic interactions
example	30
relationSource	inheritsfrom
source	https://ecologicaltraitdata.github.io/TraitDataList/#Life_span_potential

Life_span_observed

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	Life_span_observed
traitName	Life_span_observed
trait_class	life-history
broaderTerm	https://ecologicaltraitdata.github.io/TraitDataList/#life-history
valueType	numeric
traitUnit	day
maxAllowedValue	9999
minAllowedValue	0
factorLevels	NA
traitDescription	time between hatching and death
Comments	length of life-span as observed under common natural conditions including negative biotic and abiotic interactions
example	60
relationSource	refines
source	http://t-sita.cesab.org/BETSI_vizInfo.jsp?trait=Lifespan

Life_span_observed_max

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	Life_span_observed_max
traitName	Life_span_observed_max
trait_class	life-history
broaderTerm	https://ecologicaltraitdata.github.io/TraitDataList/#Life_span_observed
valueType	numeric
traitUnit	day
maxAllowedValue	9999
minAllowedValue	0
factorLevels	NA
traitDescription	maximum observed time between hatching and death
Comments	length of life-span as observed under common natural conditions including negative biotic and abiotic interactions
example	90
relationSource	inheritsfrom
source	https://ecologicaltraitdata.github.io/TraitDataList/#Life_span_observed

Life_span_observed_min

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	Life_span_observed_min
traitName	Life_span_observed_min
trait_class	life-history
broaderTerm	https://ecologicaltraitdata.github.io/TraitDataList/#Life_span_observed
valueType	numeric
traitUnit	day
maxAllowedValue	9999
minAllowedValue	0
factorLevels	NA
traitDescription	minimum observed time between hatching and death
Comments	length of life-span as observed under common natural conditions including negative biotic and abiotic interactions
example	15
relationSource	inheritsfrom
source	https://ecologicaltraitdata.github.io/TraitDataList/#Life_span_observed

Development

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	Development
traitName	Development
trait_class	life-history
broaderTerm	https://ecologicaltraitdata.github.io/TraitDataList/#life-history
valueType	character
traitUnit	NA
maxAllowedValue	NA
minAllowedValue	NA
factorLevels	NA
traitDescription	any progressive production of the phenotypic characteristics of an organism
Comments
example	NA
relationSource	equals
source	http://t-sita.cesab.org/BETSI_vizInfo.jsp?trait=Development

Age_at_maturity

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	Age_at_maturity
traitName	Age_at_maturity
trait_class	life-history
broaderTerm	https://ecologicaltraitdata.github.io/TraitDataList/#Development
valueType	numeric
traitUnit	day
maxAllowedValue	9999
minAllowedValue	0
factorLevels	NA
traitDescription	the age in months at which an individual’s physiological development is complete
Comments
example	0.5
relationSource	refines
source	http://t-sita.cesab.org/BETSI_vizInfo.jsp?trait=Maturation_time

Age_at_maturity_max

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	Age_at_maturity_max
traitName	Age_at_maturity_max
trait_class	life-history
broaderTerm	https://ecologicaltraitdata.github.io/TraitDataList/#Age_at_maturity
valueType	numeric
traitUnit	day
maxAllowedValue	9999
minAllowedValue	0
factorLevels	NA
traitDescription	maximum observed age at which an individual’s physiological development is complete
Comments
example	1
relationSource	inheritsfrom
source	https://ecologicaltraitdata.github.io/TraitDataList/#Age_at_maturity

Age_at_maturity_min

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	Age_at_maturity_min
traitName	Age_at_maturity_min
trait_class	life-history
broaderTerm	https://ecologicaltraitdata.github.io/TraitDataList/#Age_at_maturity
valueType	numeric
traitUnit	day
maxAllowedValue	9999
minAllowedValue	0
factorLevels	NA
traitDescription	minimum observed age at which an individual’s physiological development is complete
Comments
example	0.25
relationSource	inheritsfrom
source	https://ecologicaltraitdata.github.io/TraitDataList/#Age_at_maturity

Age_at_reproduction

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	Age_at_reproduction
traitName	Age_at_reproduction
trait_class	life-history
broaderTerm	https://ecologicaltraitdata.github.io/TraitDataList/#Development
valueType	numeric
traitUnit	day
maxAllowedValue	9999
minAllowedValue	0
factorLevels	NA
traitDescription	the age in months at which an individual starts reproducing
Comments
example	0.5
relationSource	inheritsfrom
source	http://t-sita.cesab.org/BETSI_vizInfo.jsp?trait=Reproduction_time

Age_at_reproduction_max

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	Age_at_reproduction_max
traitName	Age_at_reproduction_max
trait_class	life-history
broaderTerm	https://ecologicaltraitdata.github.io/TraitDataList/#Age_at_reproduction
valueType	numeric
traitUnit	day
maxAllowedValue	9999
minAllowedValue	0
factorLevels	NA
traitDescription	maximum observed age at which an individual starts reproducing
Comments
example	1
relationSource	inheritsfrom
source	https://ecologicaltraitdata.github.io/TraitDataList/#Age_at_reproduction

Age_at_reproduction_min

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	Age_at_reproduction_min
traitName	Age_at_reproduction_min
trait_class	life-history
broaderTerm	https://ecologicaltraitdata.github.io/TraitDataList/#Age_at_reproduction
valueType	numeric
traitUnit	day
maxAllowedValue	9999
minAllowedValue	0
factorLevels	NA
traitDescription	minimum observed age at which an individual starts reproducing
Comments
example	0.25
relationSource	inheritsfrom
source	https://ecologicaltraitdata.github.io/TraitDataList/#Age_at_reproduction

Morphological traits

Reproduction_morphology

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	Reproduction_morphology
traitName	Reproduction_morphology
trait_class	morphology
broaderTerm	https://ecologicaltraitdata.github.io/TraitDataList/#morphology
valueType	character
traitUnit	NA
maxAllowedValue	NA
minAllowedValue	NA
factorLevels	NA
traitDescription	any morphological structure which serves a specialized function for reproduction
Comments
example	NA
relationSource	equals
source	http://t-sita.cesab.org/BETSI_vizInfo.jsp?trait=Reproduction

Sperm_storage

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	Sperm_storage
traitName	Sperm_storage
trait_class	morphology
broaderTerm	https://ecologicaltraitdata.github.io/TraitDataList/#Reproduction_morphology
valueType	logical
traitUnit	NA
maxAllowedValue	NA
minAllowedValue	NA
factorLevels	NA
traitDescription	presence of a sperm storage organ
Comments	A collection of tissues joined in structural unit to keep some male gametes for future use
example	TRUE
relationSource	equals
source	http://t-sita.cesab.org/BETSI_vizInfo.jsp?trait=Sperm_storage_organ

Nr_Ovariols

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	Nr_Ovariols
traitName	Nr_Ovariols
trait_class	morphology
broaderTerm	https://ecologicaltraitdata.github.io/TraitDataList/#Reproduction_morphology
valueType	numeric
traitUnit	count
maxAllowedValue	999
minAllowedValue	1
factorLevels	NA
traitDescription	number of ovariols per individual
Comments
example	5
relationSource
source

Nr_Ovariols_max

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	Nr_Ovariols_max
traitName	Nr_Ovariols_max
trait_class	morphology
broaderTerm	https://ecologicaltraitdata.github.io/TraitDataList/#Nr_Ovariols
valueType	numeric
traitUnit	count
maxAllowedValue	999
minAllowedValue	1
factorLevels	NA
traitDescription	maximum observed number of ovariols per individual
Comments
example	5
relationSource	inheritsfrom
source	https://ecologicaltraitdata.github.io/TraitDataList/#Nr_Ovariols

Nr_Ovariols_min

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	Nr_Ovariols_min
traitName	Nr_Ovariols_min
trait_class	morphology
broaderTerm	https://ecologicaltraitdata.github.io/TraitDataList/#Nr_Ovariols
valueType	numeric
traitUnit	count
maxAllowedValue	999
minAllowedValue	1
factorLevels	NA
traitDescription	minimum observed number of ovariols per individual
Comments
example	5
relationSource	inheritsfrom
source	https://ecologicaltraitdata.github.io/TraitDataList/#Nr_Ovariols

Body_shape

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	Body_shape
traitName	Body_shape
trait_class	morphology
broaderTerm	https://ecologicaltraitdata.github.io/TraitDataList/#morphology
valueType	factor
traitUnit	NA
maxAllowedValue	NA
minAllowedValue	NA
factorLevels	spherical/cylindrical/ovoid/trapezoidal
traitDescription	Geometrical description of the part of that space occupied by the physical structure of an organism
Comments
example	spherical
relationSource	equals
source	http://t-sita.cesab.org/BETSI_vizInfo.jsp?trait=Body_shape

Body_size

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	Body_size
traitName	Body_size
trait_class	morphology
broaderTerm	https://ecologicaltraitdata.github.io/TraitDataList/#morphology
valueType	factor
traitUnit	NA
maxAllowedValue	NA
minAllowedValue	NA
factorLevels	<1mm/<1cm/<10cm/<100m
traitDescription	general size class
Comments
example	<1cm
relationSource	refines
source	http://t-sita.cesab.org/BETSI_vizInfo.jsp?trait=Body_dimension

Body_length

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	Body_length
traitName	Body_length
trait_class	morphology
broaderTerm	https://ecologicaltraitdata.github.io/TraitDataList/#Body_size
valueType	numeric
traitUnit	cm
maxAllowedValue	999
minAllowedValue	0
factorLevels	NA
traitDescription	longest dimension of the physical structure of organisms
Comments
example	0.75
relationSource	equals
source	http://t-sita.cesab.org/BETSI_vizInfo.jsp?trait=Body_length

Body_length_max

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	Body_length_max
traitName	Body_length_max
trait_class	morphology
broaderTerm	https://ecologicaltraitdata.github.io/TraitDataList/#Body_length
valueType	numeric
traitUnit	cm
maxAllowedValue	999
minAllowedValue	0
factorLevels	NA
traitDescription	maximum documented body length
Comments
example	1.25
relationSource	inheritsfrom
source	https://ecologicaltraitdata.github.io/TraitDataList/#Body_length

Body_length_min

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	Body_length_min
traitName	Body_length_min
trait_class	morphology
broaderTerm	https://ecologicaltraitdata.github.io/TraitDataList/#Body_length
valueType	numeric
traitUnit	cm
maxAllowedValue	999
minAllowedValue	0
factorLevels	NA
traitDescription	minimum documented body length
Comments
example	0.25
relationSource	inheritsfrom
source	https://ecologicaltraitdata.github.io/TraitDataList/#Body_length

Pronotum_length

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	Pronotum_length
traitName	Pronotum_length
trait_class	morphology
broaderTerm	https://ecologicaltraitdata.github.io/TraitDataList/#Body_length
valueType	numeric
traitUnit	cm
maxAllowedValue	999
minAllowedValue	0
factorLevels	NA
traitDescription	longest dimension of the pronotum
Comments
example	0.75
relationSource	refines
source	http://t-sita.cesab.org/BETSI_vizInfo.jsp?trait=Body_length

Pronotum_length_max

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	Pronotum_length_max
traitName	Pronotum_length_max
trait_class	morphology
broaderTerm	https://ecologicaltraitdata.github.io/TraitDataList/#Pronotum_length
valueType	numeric
traitUnit	cm
maxAllowedValue	999
minAllowedValue	0
factorLevels	NA
traitDescription	maximum documented pronotum length
Comments
example	1.25
relationSource	inheritsfrom
source	https://ecologicaltraitdata.github.io/TraitDataList/#Pronotum_length

Pronotum_length_min

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	Pronotum_length_min
traitName	Pronotum_length_min
trait_class	morphology
broaderTerm	https://ecologicaltraitdata.github.io/TraitDataList/#Pronotum_length
valueType	numeric
traitUnit	cm
maxAllowedValue	999
minAllowedValue	0
factorLevels	NA
traitDescription	minimum documented pronotum length
Comments
example	0.5
relationSource	inheritsfrom
source	https://ecologicaltraitdata.github.io/TraitDataList/#Pronotum_length

Thorax_length

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	Thorax_length
traitName	Thorax_length
trait_class	morphology
broaderTerm	https://ecologicaltraitdata.github.io/TraitDataList/#Body_length
valueType	numeric
traitUnit	cm
maxAllowedValue	999
minAllowedValue	0
factorLevels	NA
traitDescription	longest dimension of the thorax
Comments
example	0.75
relationSource	refines
source	http://t-sita.cesab.org/BETSI_vizInfo.jsp?trait=Body_length

Thorax_length_max

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	Thorax_length_max
traitName	Thorax_length_max
trait_class	morphology
broaderTerm	https://ecologicaltraitdata.github.io/TraitDataList/#Thorax_length
valueType	numeric
traitUnit	cm
maxAllowedValue	999
minAllowedValue	0
factorLevels	NA
traitDescription	maximum documented thorax length
Comments
example	1.25
relationSource	inheritsfrom
source	https://ecologicaltraitdata.github.io/TraitDataList/#Thorax_length

Thorax_length_min

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	Thorax_length_min
traitName	Thorax_length_min
trait_class	morphology
broaderTerm	https://ecologicaltraitdata.github.io/TraitDataList/#Thorax_length
valueType	numeric
traitUnit	cm
maxAllowedValue	999
minAllowedValue	0
factorLevels	NA
traitDescription	minimum documented thorax length
Comments
example	0.5
relationSource	inheritsfrom
source	https://ecologicaltraitdata.github.io/TraitDataList/#Thorax_length

Head_length

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	Head_length
traitName	Head_length
trait_class	morphology
broaderTerm	https://ecologicaltraitdata.github.io/TraitDataList/#Body_length
valueType	numeric
traitUnit	cm
maxAllowedValue	999
minAllowedValue	0
factorLevels	NA
traitDescription	longest dimension of the head
Comments
example	0.75
relationSource	refines
source	http://t-sita.cesab.org/BETSI_vizInfo.jsp?trait=Body_length

Head_length_max

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	Head_length_max
traitName	Head_length_max
trait_class	morphology
broaderTerm	https://ecologicaltraitdata.github.io/TraitDataList/#Head_length
valueType	numeric
traitUnit	cm
maxAllowedValue	999
minAllowedValue	0
factorLevels	NA
traitDescription	maximum documented head length
Comments
example	1.25
relationSource	inheritsfrom
source	https://ecologicaltraitdata.github.io/TraitDataList/#Head_length

Head_length_min

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	Head_length_min
traitName	Head_length_min
trait_class	morphology
broaderTerm	https://ecologicaltraitdata.github.io/TraitDataList/#Head_length
valueType	numeric
traitUnit	cm
maxAllowedValue	999
minAllowedValue	0
factorLevels	NA
traitDescription	minimum documented head length
Comments
example	0.5
relationSource	inheritsfrom
source	https://ecologicaltraitdata.github.io/TraitDataList/#Head_length

Body_width

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	Body_width
traitName	Body_width
trait_class	morphology
broaderTerm	https://ecologicaltraitdata.github.io/TraitDataList/#Body_size
valueType	numeric
traitUnit	cm
maxAllowedValue	999
minAllowedValue	0
factorLevels	NA
traitDescription	second longest dimension of the physical structure of organisms
Comments
example	0.25
relationSource	equals
source	http://t-sita.cesab.org/BETSI_vizInfo.jsp?trait=Body_width

Body_width_max

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	Body_width_max
traitName	Body_width_max
trait_class	morphology
broaderTerm	https://ecologicaltraitdata.github.io/TraitDataList/#Body_width
valueType	numeric
traitUnit	cm
maxAllowedValue	999
minAllowedValue	0
factorLevels	NA
traitDescription	maximum documented body width
Comments
example	0.5
relationSource	inheritsfrom
source	https://ecologicaltraitdata.github.io/TraitDataList/#Body_width

Body_width_min

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	Body_width_min
traitName	Body_width_min
trait_class	morphology
broaderTerm	https://ecologicaltraitdata.github.io/TraitDataList/#Body_width
valueType	numeric
traitUnit	cm
maxAllowedValue	999
minAllowedValue	0
factorLevels	NA
traitDescription	minimum documented body width
Comments
example	0.25
relationSource	inheritsfrom
source	https://ecologicaltraitdata.github.io/TraitDataList/#Body_width

Pronotum_width

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	Pronotum_width
traitName	Pronotum_width
trait_class	morphology
broaderTerm	https://ecologicaltraitdata.github.io/TraitDataList/#Body_width
valueType	numeric
traitUnit	cm
maxAllowedValue	999
minAllowedValue	0
factorLevels	NA
traitDescription	second longest dimension of the pronotum
Comments
example	0.5
relationSource	refines
source	http://t-sita.cesab.org/BETSI_vizInfo.jsp?trait=Body_width

Pronotum_width_max

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	Pronotum_width_max
traitName	Pronotum_width_max
trait_class	morphology
broaderTerm	https://ecologicaltraitdata.github.io/TraitDataList/#Pronotum_width
valueType	numeric
traitUnit	cm
maxAllowedValue	999
minAllowedValue	0
factorLevels	NA
traitDescription	maximum documented pronotum width
Comments
example	1.25
relationSource	inheritsfrom
source	https://ecologicaltraitdata.github.io/TraitDataList/#Pronotum_width

Pronotum_width_min

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	Pronotum_width_min
traitName	Pronotum_width_min
trait_class	morphology
broaderTerm	https://ecologicaltraitdata.github.io/TraitDataList/#Pronotum_width
valueType	numeric
traitUnit	cm
maxAllowedValue	999
minAllowedValue	0
factorLevels	NA
traitDescription	minimum documented pronotum width
Comments
example	0.5
relationSource	inheritsfrom
source	https://ecologicaltraitdata.github.io/TraitDataList/#Pronotum_width

Intertegular_span

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	Intertegular_span
traitName	Intertegular_span
trait_class	morphology
broaderTerm	https://ecologicaltraitdata.github.io/TraitDataList/#Body_width
valueType	numeric
traitUnit	cm
maxAllowedValue	999
minAllowedValue	0
factorLevels	NA
traitDescription	distance between the two tegulae
Comments
example	0.5
relationSource	refines
source	http://t-sita.cesab.org/BETSI_vizInfo.jsp?trait=Body_width

Intertegular_span_max

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	Intertegular_span_max
traitName	Intertegular_span_max
trait_class	morphology
broaderTerm	https://ecologicaltraitdata.github.io/TraitDataList/#Intertegular_span
valueType	numeric
traitUnit	cm
maxAllowedValue	999
minAllowedValue	0
factorLevels	NA
traitDescription	maximum documented distance between the two tegulae
Comments
example	0.55
relationSource	inheritsfrom
source	https://ecologicaltraitdata.github.io/TraitDataList/#Intertegular_span

Intertegular_span_min

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	Intertegular_span_min
traitName	Intertegular_span_min
trait_class	morphology
broaderTerm	https://ecologicaltraitdata.github.io/TraitDataList/#Intertegular_span
valueType	numeric
traitUnit	cm
maxAllowedValue	999
minAllowedValue	0
factorLevels	NA
traitDescription	minimum documented distance between the two tegulae
Comments
example	0.45
relationSource	inheritsfrom
source	https://ecologicaltraitdata.github.io/TraitDataList/#Intertegular_span

Thorax_width

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	Thorax_width
traitName	Thorax_width
trait_class	morphology
broaderTerm	https://ecologicaltraitdata.github.io/TraitDataList/#Body_width
valueType	numeric
traitUnit	cm
maxAllowedValue	999
minAllowedValue	0
factorLevels	NA
traitDescription	second longest dimension of the thorax
Comments
example	0.5
relationSource	refines
source	http://t-sita.cesab.org/BETSI_vizInfo.jsp?trait=Body_width

Thorax_width_max

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	Thorax_width_max
traitName	Thorax_width_max
trait_class	morphology
broaderTerm	https://ecologicaltraitdata.github.io/TraitDataList/#Thorax_width
valueType	numeric
traitUnit	cm
maxAllowedValue	999
minAllowedValue	0
factorLevels	NA
traitDescription	maximum documented thorax width
Comments
example	1.25
relationSource	inheritsfrom
source	https://ecologicaltraitdata.github.io/TraitDataList/#Thorax_width

Thorax_width_min

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	Thorax_width_min
traitName	Thorax_width_min
trait_class	morphology
broaderTerm	https://ecologicaltraitdata.github.io/TraitDataList/#Thorax_width
valueType	numeric
traitUnit	cm
maxAllowedValue	999
minAllowedValue	0
factorLevels	NA
traitDescription	minimum documented thorax width
Comments
example	0.5
relationSource	inheritsfrom
source	https://ecologicaltraitdata.github.io/TraitDataList/#Thorax_width

Head_width

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	Head_width
traitName	Head_width
trait_class	morphology
broaderTerm	https://ecologicaltraitdata.github.io/TraitDataList/#Body_width
valueType	numeric
traitUnit	cm
maxAllowedValue	999
minAllowedValue	0
factorLevels	NA
traitDescription	second longest dimension of the head
Comments
example	0.5
relationSource	refines
source	http://t-sita.cesab.org/BETSI_vizInfo.jsp?trait=Body_width

Head_width_max

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	Head_width_max
traitName	Head_width_max
trait_class	morphology
broaderTerm	https://ecologicaltraitdata.github.io/TraitDataList/#Head_width
valueType	numeric
traitUnit	cm
maxAllowedValue	999
minAllowedValue	0
factorLevels	NA
traitDescription	maximum documented head width
Comments
example	1.25
relationSource	inheritsfrom
source	https://ecologicaltraitdata.github.io/TraitDataList/#Head_width

Head_width_min

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	Head_width_min
traitName	Head_width_min
trait_class	morphology
broaderTerm	https://ecologicaltraitdata.github.io/TraitDataList/#Head_width
valueType	numeric
traitUnit	cm
maxAllowedValue	999
minAllowedValue	0
factorLevels	NA
traitDescription	minimum documented head width
Comments
example	0.5
relationSource	inheritsfrom
source	https://ecologicaltraitdata.github.io/TraitDataList/#Head_width

Body_height

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	Body_height
traitName	Body_height
trait_class	morphology
broaderTerm	https://ecologicaltraitdata.github.io/TraitDataList/#Body_size
valueType	numeric
traitUnit	cm
maxAllowedValue	999
minAllowedValue	0
factorLevels	NA
traitDescription	third longest dimension of the physical structure of organisms
Comments
example	0.25
relationSource	inheritsfrom
source	http://t-sita.cesab.org/BETSI_vizInfo.jsp?trait=Body_width

Body_height_max

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	Body_height_max
traitName	Body_height_max
trait_class	morphology
broaderTerm	https://ecologicaltraitdata.github.io/TraitDataList/#Body_height
valueType	numeric
traitUnit	cm
maxAllowedValue	999
minAllowedValue	0
factorLevels	NA
traitDescription	maximum documented body height
Comments
example	0.5
relationSource	inheritsfrom
source	https://ecologicaltraitdata.github.io/TraitDataList/#Body_height

Body_height_min

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	Body_height_min
traitName	Body_height_min
trait_class	morphology
broaderTerm	https://ecologicaltraitdata.github.io/TraitDataList/#Body_height
valueType	numeric
traitUnit	cm
maxAllowedValue	999
minAllowedValue	0
factorLevels	NA
traitDescription	minimum documented body height
Comments
example	0.15
relationSource	inheritsfrom
source	https://ecologicaltraitdata.github.io/TraitDataList/#Body_height

Pronotum_height

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	Pronotum_height
traitName	Pronotum_height
trait_class	morphology
broaderTerm	https://ecologicaltraitdata.github.io/TraitDataList/#Body_height
valueType	numeric
traitUnit	cm
maxAllowedValue	999
minAllowedValue	0
factorLevels	Na
traitDescription	third longest dimension of the pronotum
Comments
example	0.25
relationSource	refines
source	http://t-sita.cesab.org/BETSI_vizInfo.jsp?trait=Body_width

Pronotum_height_max

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	Pronotum_height_max
traitName	Pronotum_height_max
trait_class	morphology
broaderTerm	https://ecologicaltraitdata.github.io/TraitDataList/#Pronotum_height
valueType	numeric
traitUnit	cm
maxAllowedValue	999
minAllowedValue	0
factorLevels	NA
traitDescription	maximum documented pronotum height
Comments
example	1.25
relationSource	inheritsfrom
source	https://ecologicaltraitdata.github.io/TraitDataList/#Pronotum_height

Pronotum_height_min

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	Pronotum_height_min
traitName	Pronotum_height_min
trait_class	morphology
broaderTerm	https://ecologicaltraitdata.github.io/TraitDataList/#Pronotum_height
valueType	numeric
traitUnit	cm
maxAllowedValue	999
minAllowedValue	0
factorLevels	NA
traitDescription	minimum documented pronotum height
Comments
example	0.5
relationSource	inheritsfrom
source	https://ecologicaltraitdata.github.io/TraitDataList/#Pronotum_height

Thorax_height

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	Thorax_height
traitName	Thorax_height
trait_class	morphology
broaderTerm	https://ecologicaltraitdata.github.io/TraitDataList/#Body_height
valueType	numeric
traitUnit	cm
maxAllowedValue	999
minAllowedValue	0
factorLevels	Na
traitDescription	third longest dimension of the thorax
Comments
example	0.25
relationSource	refines
source	http://t-sita.cesab.org/BETSI_vizInfo.jsp?trait=Body_width

Thorax_height_max

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	Thorax_height_max
traitName	Thorax_height_max
trait_class	morphology
broaderTerm	https://ecologicaltraitdata.github.io/TraitDataList/#Thorax_height
valueType	numeric
traitUnit	cm
maxAllowedValue	999
minAllowedValue	0
factorLevels	NA
traitDescription	maximum documented thorax height
Comments
example	1.25
relationSource	inheritsfrom
source	https://ecologicaltraitdata.github.io/TraitDataList/#Thorax_height

Thorax_height_min

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	Thorax_height_min
traitName	Thorax_height_min
trait_class	morphology
broaderTerm	https://ecologicaltraitdata.github.io/TraitDataList/#Thorax_height
valueType	numeric
traitUnit	cm
maxAllowedValue	999
minAllowedValue	0
factorLevels	NA
traitDescription	minimum documented thorax height
Comments
example	0.5
relationSource	inheritsfrom
source	https://ecologicaltraitdata.github.io/TraitDataList/#Thorax_height

Head_height

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	Head_height
traitName	Head_height
trait_class	morphology
broaderTerm	https://ecologicaltraitdata.github.io/TraitDataList/#Body_height
valueType	numeric
traitUnit	cm
maxAllowedValue	999
minAllowedValue	0
factorLevels	Na
traitDescription	third longest dimension of the head
Comments
example	0.25
relationSource	refines
source	http://t-sita.cesab.org/BETSI_vizInfo.jsp?trait=Body_width

Head_height_max

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	Head_height_max
traitName	Head_height_max
trait_class	morphology
broaderTerm	https://ecologicaltraitdata.github.io/TraitDataList/#Head_height
valueType	numeric
traitUnit	cm
maxAllowedValue	999
minAllowedValue	0
factorLevels	NA
traitDescription	maximum documented head height
Comments
example	1.25
relationSource	inheritsfrom
source	https://ecologicaltraitdata.github.io/TraitDataList/#Head_height

Head_height_min

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	Head_height_min
traitName	Head_height_min
trait_class	morphology
broaderTerm	https://ecologicaltraitdata.github.io/TraitDataList/#Head_height
valueType	numeric
traitUnit	cm
maxAllowedValue	999
minAllowedValue	0
factorLevels	NA
traitDescription	minimum documented head height
Comments
example	0.5
relationSource	inheritsfrom
source	https://ecologicaltraitdata.github.io/TraitDataList/#Head_height

Body_mass_live

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	Body_mass_live
traitName	Body_mass_live
trait_class	morphology
broaderTerm	https://ecologicaltraitdata.github.io/TraitDataList/#Body_size
valueType	numeric
traitUnit	g
maxAllowedValue	999
minAllowedValue	0
factorLevels	NA
traitDescription	body weight measured on a living individual
Comments
example	20
relationSource	refines
source	http://t-sita.cesab.org/BETSI_vizInfo.jsp?trait=Fresh_body_weight

Body_mass_live_max

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	Body_mass_live_max
traitName	Body_mass_live_max
trait_class	morphology
broaderTerm	https://ecologicaltraitdata.github.io/TraitDataList/#Body_mass_live
valueType	numeric
traitUnit	g
maxAllowedValue	999
minAllowedValue	0
factorLevels	NA
traitDescription	maximum body weight measured on a living individual
Comments
example	25
relationSource	inheritsfrom
source	https://ecologicaltraitdata.github.io/TraitDataList/#Body_mass_live

Body_mass_live_min

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	Body_mass_live_min
traitName	Body_mass_live_min
trait_class	morphology
broaderTerm	https://ecologicaltraitdata.github.io/TraitDataList/#Body_mass_live
valueType	numeric
traitUnit	g
maxAllowedValue	999
minAllowedValue	0
factorLevels	NA
traitDescription	minimum body weight measured on a living individual
Comments
example	15
relationSource	inheritsfrom
source	https://ecologicaltraitdata.github.io/TraitDataList/#Body_mass_live

Body_mass_fresh

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	Body_mass_fresh
traitName	Body_mass_fresh
trait_class	morphology
broaderTerm	https://ecologicaltraitdata.github.io/TraitDataList/#Body_size
valueType	numeric
traitUnit	g
maxAllowedValue	999
minAllowedValue	0
factorLevels	NA
traitDescription	body weight measured on a freshly dead individual
Comments
example	20
relationSource	equals
source	http://t-sita.cesab.org/BETSI_vizInfo.jsp?trait=Fresh_body_weight

Body_mass_fresh_max

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	Body_mass_fresh_max
traitName	Body_mass_fresh_max
trait_class	morphology
broaderTerm	https://ecologicaltraitdata.github.io/TraitDataList/#Body_mass_fresh
valueType	numeric
traitUnit	g
maxAllowedValue	999
minAllowedValue	0
factorLevels	NA
traitDescription	maximum body weight measured on a freshly dead individual
Comments
example	25
relationSource	inheritsfrom
source	https://ecologicaltraitdata.github.io/TraitDataList/#Body_mass_fresh

Body_mass_fresh_min

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	Body_mass_fresh_min
traitName	Body_mass_fresh_min
trait_class	morphology
broaderTerm	https://ecologicaltraitdata.github.io/TraitDataList/#Body_mass_fresh
valueType	numeric
traitUnit	g
maxAllowedValue	999
minAllowedValue	0
factorLevels	NA
traitDescription	minimum body weight measured on a freshly dead individual
Comments
example	15
relationSource	inheritsfrom
source	https://ecologicaltraitdata.github.io/TraitDataList/#Body_mass_fresh

Body_mass_dried

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	Body_mass_dried
traitName	Body_mass_dried
trait_class	morphology
broaderTerm	https://ecologicaltraitdata.github.io/TraitDataList/#Body_size
valueType	numeric
traitUnit	g
maxAllowedValue	999
minAllowedValue	0
factorLevels	NA
traitDescription	body weight measured on a dried individual
Comments
example	10
relationSource	equals
source	http://t-sita.cesab.org/BETSI_vizInfo.jsp?trait=Dry_body_weight

Body_mass_dried_max

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	Body_mass_dried_max
traitName	Body_mass_dried_max
trait_class	morphology
broaderTerm	https://ecologicaltraitdata.github.io/TraitDataList/#Body_mass_dried
valueType	numeric
traitUnit	g
maxAllowedValue	999
minAllowedValue	0
factorLevels	NA
traitDescription	maximum body weight measured on a dried individual
Comments
example	15
relationSource	inheritsfrom
source	https://ecologicaltraitdata.github.io/TraitDataList/#Body_mass_dried

Body_mass_dried_min

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	Body_mass_dried_min
traitName	Body_mass_dried_min
trait_class	morphology
broaderTerm	https://ecologicaltraitdata.github.io/TraitDataList/#Body_mass_dried
valueType	numeric
traitUnit	g
maxAllowedValue	999
minAllowedValue	0
factorLevels	NA
traitDescription	minimum body weight measured on a dried individual
Comments
example	5
relationSource	inheritsfrom
source	https://ecologicaltraitdata.github.io/TraitDataList/#Body_mass_dried

Body_volume

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	Body_volume
traitName	Body_volume
trait_class	morphology
broaderTerm	https://ecologicaltraitdata.github.io/TraitDataList/#Body_size
valueType	numeric
traitUnit	cm³
maxAllowedValue	9999
minAllowedValue	0
factorLevels	NA
traitDescription	body volume
Comments	usually calculated from measures of length/width/height
example	0.75
relationSource	refines
source	http://t-sita.cesab.org/BETSI_vizInfo.jsp?trait=Body_dimension

Body_volume_max

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	Body_volume_max
traitName	Body_volume_max
trait_class	morphology
broaderTerm	https://ecologicaltraitdata.github.io/TraitDataList/#Body_volume
valueType	numeric
traitUnit	cm³
maxAllowedValue	9999
minAllowedValue	0
factorLevels	NA
traitDescription	maximum documented body volume
Comments	usually calculated from measures of length/width/height
example	1
relationSource	inheritsfrom
source	https://ecologicaltraitdata.github.io/TraitDataList/#Body_volume

Body_volume_min

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	Body_volume_min
traitName	Body_volume_min
trait_class	morphology
broaderTerm	https://ecologicaltraitdata.github.io/TraitDataList/#Body_volume
valueType	numeric
traitUnit	cm³
maxAllowedValue	9999
minAllowedValue	0
factorLevels	NA
traitDescription	minimum documented body volume
Comments	usually calculated from measures of length/width/height
example	0.56
relationSource	inheritsfrom
source	https://ecologicaltraitdata.github.io/TraitDataList/#Body_volume

Length_width_ratio

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	Length_width_ratio
traitName	Length_width_ratio
trait_class	morphology
broaderTerm	https://ecologicaltraitdata.github.io/TraitDataList/#Body_size
valueType	numeric
traitUnit	ratio
maxAllowedValue	999
minAllowedValue	0
factorLevels	NA
traitDescription	ratio between body length and body width
Comments
example	2
relationSource	refines
source	http://t-sita.cesab.org/BETSI_vizInfo.jsp?trait=Body_dimension

Length_width_ratio_max

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	Length_width_ratio_max
traitName	Length_width_ratio_max
trait_class	morphology
broaderTerm	https://ecologicaltraitdata.github.io/TraitDataList/#Length_width_ratio
valueType	numeric
traitUnit	ratio
maxAllowedValue	999
minAllowedValue	0
factorLevels	NA
traitDescription	maximum documented ratio between body length and body width
Comments
example	2.5
relationSource	inheritsfrom
source	https://ecologicaltraitdata.github.io/TraitDataList/#Length_width_ratio

Length_width_ratio_min

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	Length_width_ratio_min
traitName	Length_width_ratio_min
trait_class	morphology
broaderTerm	https://ecologicaltraitdata.github.io/TraitDataList/#Length_width_ratio
valueType	numeric
traitUnit	ratio
maxAllowedValue	999
minAllowedValue	0
factorLevels	NA
traitDescription	minimum documented ratio between body length and body width
Comments
example	1.75
relationSource	inheritsfrom
source	https://ecologicaltraitdata.github.io/TraitDataList/#Length_width_ratio

Length_width_ratio_pronotum

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	Length_width_ratio_pronotum
traitName	Length_width_ratio_pronotum
trait_class	morphology
broaderTerm	https://ecologicaltraitdata.github.io/TraitDataList/#Length_width_ratio
valueType	numeric
traitUnit	ratio
maxAllowedValue	999
minAllowedValue	0
factorLevels	NA
traitDescription	ratio between pronotum length and width
Comments
example	2
relationSource	inheritsfrom
source	https://ecologicaltraitdata.github.io/TraitDataList/#Length_width_ratio

Length_width_ratio_pronotum_max

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	Length_width_ratio_pronotum_max
traitName	Length_width_ratio_pronotum_max
trait_class	morphology
broaderTerm	https://ecologicaltraitdata.github.io/TraitDataList/#Length_width_ratio_pronotum
valueType	numeric
traitUnit	ratio
maxAllowedValue	999
minAllowedValue	0
factorLevels	NA
traitDescription	maximum documented ratio between pronotum length and width
Comments
example	2.5
relationSource	inheritsfrom
source	https://ecologicaltraitdata.github.io/TraitDataList/#Length_width_ratio_pronotum

Length_width_ratio_pronotum_min

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	Length_width_ratio_pronotum_min
traitName	Length_width_ratio_pronotum_min
trait_class	morphology
broaderTerm	https://ecologicaltraitdata.github.io/TraitDataList/#Length_width_ratio_pronotum
valueType	numeric
traitUnit	ratio
maxAllowedValue	999
minAllowedValue	0
factorLevels	NA
traitDescription	minimum documented ratio between pronotum length and width
Comments
example	1.75
relationSource	inheritsfrom
source	https://ecologicaltraitdata.github.io/TraitDataList/#Length_width_ratio_pronotum

Length_width_ratio_thorax

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	Length_width_ratio_thorax
traitName	Length_width_ratio_thorax
trait_class	morphology
broaderTerm	https://ecologicaltraitdata.github.io/TraitDataList/#Length_width_ratio
valueType	numeric
traitUnit	ratio
maxAllowedValue	999
minAllowedValue	0
factorLevels	NA
traitDescription	ratio between thorax length and width
Comments
example	2
relationSource	inheritsfrom
source	https://ecologicaltraitdata.github.io/TraitDataList/#Length_width_ratio

Length_width_ratio_thorax_max

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	Length_width_ratio_thorax_max
traitName	Length_width_ratio_thorax_max
trait_class	morphology
broaderTerm	https://ecologicaltraitdata.github.io/TraitDataList/#Length_width_ratio_thorax
valueType	numeric
traitUnit	ratio
maxAllowedValue	999
minAllowedValue	0
factorLevels	NA
traitDescription	maximum documented ratio between thorax length and width
Comments
example	2.5
relationSource	inheritsfrom
source	https://ecologicaltraitdata.github.io/TraitDataList/#Length_width_ratio_thorax

Length_width_ratio_thorax_min

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	Length_width_ratio_thorax_min
traitName	Length_width_ratio_thorax_min
trait_class	morphology
broaderTerm	https://ecologicaltraitdata.github.io/TraitDataList/#Length_width_ratio_thorax
valueType	numeric
traitUnit	ratio
maxAllowedValue	999
minAllowedValue	0
factorLevels	NA
traitDescription	minimum documented ratio between thorax length and width
Comments
example	1.75
relationSource	inheritsfrom
source	https://ecologicaltraitdata.github.io/TraitDataList/#Length_width_ratio_thorax

Length_width_ratio_head

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	Length_width_ratio_head
traitName	Length_width_ratio_head
trait_class	morphology
broaderTerm	https://ecologicaltraitdata.github.io/TraitDataList/#Length_width_ratio
valueType	numeric
traitUnit	ratio
maxAllowedValue	999
minAllowedValue	0
factorLevels	NA
traitDescription	ratio between head length and width
Comments
example	2
relationSource	inheritsfrom
source	https://ecologicaltraitdata.github.io/TraitDataList/#Length_width_ratio

Length_width_ratio_head_max

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	Length_width_ratio_head_max
traitName	Length_width_ratio_head_max
trait_class	morphology
broaderTerm	https://ecologicaltraitdata.github.io/TraitDataList/#Length_width_ratio_head
valueType	numeric
traitUnit	ratio
maxAllowedValue	999
minAllowedValue	0
factorLevels	NA
traitDescription	maximum documented ratio between head length and width
Comments
example	2.5
relationSource	inheritsfrom
source	https://ecologicaltraitdata.github.io/TraitDataList/#Length_width_ratio_head

Length_width_ratio_head_min

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	Length_width_ratio_head_min
traitName	Length_width_ratio_head_min
trait_class	morphology
broaderTerm	https://ecologicaltraitdata.github.io/TraitDataList/#Length_width_ratio_head
valueType	numeric
traitUnit	ratio
maxAllowedValue	999
minAllowedValue	0
factorLevels	NA
traitDescription	minimum documented ratio between head length and width
Comments
example	1.75
relationSource	inheritsfrom
source	https://ecologicaltraitdata.github.io/TraitDataList/#Length_width_ratio_head

Nutrition_morphology

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	Nutrition_morphology
traitName	Nutrition_morphology
trait_class	morphology
broaderTerm	https://ecologicaltraitdata.github.io/TraitDataList/#morphology
valueType	character
traitUnit	NA
maxAllowedValue	NA
minAllowedValue	NA
factorLevels	NA
traitDescription	any morphological structure which serves a specialized function for nutrition
Comments
example	NA
relationSource	equals
source	http://t-sita.cesab.org/BETSI_vizInfo.jsp?trait=Nutrition

Mouthpart_type

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	Mouthpart_type
traitName	Mouthpart_type
trait_class	morphology
broaderTerm	https://ecologicaltraitdata.github.io/TraitDataList/#Nutrition_morphology
valueType	factor
traitUnit	NA
maxAllowedValue	NA
minAllowedValue	NA
factorLevels	absent/sucking/swallowing/chewing/piercing-sucking
traitDescription	category of mouthpart following the feeding strategy
Comments	Any appendage adapted to a specific way of ingesting which shares similar characteristics and forms a smaller division of a larger set
example	sucking
relationSource	equals
source	http://t-sita.cesab.org/BETSI_vizInfo.jsp?trait=Mouthpart_type

Mandible_width

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	Mandible_width
traitName	Mandible_width
trait_class	morphology
broaderTerm	https://ecologicaltraitdata.github.io/TraitDataList/#Nutrition_morphology
valueType	numeric
traitUnit	mm
maxAllowedValue	999
minAllowedValue	0
factorLevels	NA
traitDescription	width of a single mandible
Comments
example	0.02
relationSource	equals
source	http://t-sita.cesab.org/BETSI_vizInfo.jsp?trait=Mandible_width

Mandible_width_max

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	Mandible_width_max
traitName	Mandible_width_max
trait_class	morphology
broaderTerm	https://ecologicaltraitdata.github.io/TraitDataList/#Mandible_width
valueType	numeric
traitUnit	mm
maxAllowedValue	999
minAllowedValue	0
factorLevels	NA
traitDescription	maximum documented width of a single mandible
Comments
example	0.05
relationSource	inheritsfrom
source	https://ecologicaltraitdata.github.io/TraitDataList/#Mandible_width

Mandible_width_min

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	Mandible_width_min
traitName	Mandible_width_min
trait_class	morphology
broaderTerm	https://ecologicaltraitdata.github.io/TraitDataList/#Mandible_width
valueType	numeric
traitUnit	mm
maxAllowedValue	999
minAllowedValue	0
factorLevels	NA
traitDescription	minimum documented width of a single mandible
Comments
example	0.01
relationSource	inheritsfrom
source	https://ecologicaltraitdata.github.io/TraitDataList/#Mandible_width

Mandible_length

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	Mandible_length
traitName	Mandible_length
trait_class	morphology
broaderTerm	https://ecologicaltraitdata.github.io/TraitDataList/#Nutrition_morphology
valueType	numeric
traitUnit	mm
maxAllowedValue	999
minAllowedValue	0
factorLevels	NA
traitDescription	length of a single mandible
Comments
example	0.02
relationSource	equals
source	http://t-sita.cesab.org/BETSI_vizInfo.jsp?trait=Mandible_length

Mandible_length_max

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	Mandible_length_max
traitName	Mandible_length_max
trait_class	morphology
broaderTerm	https://ecologicaltraitdata.github.io/TraitDataList/#Mandible_length
valueType	numeric
traitUnit	mm
maxAllowedValue	999
minAllowedValue	0
factorLevels	NA
traitDescription	maximum documented length of a single mandible
Comments
example	0.05
relationSource	inheritsfrom
source	https://ecologicaltraitdata.github.io/TraitDataList/#Mandible_length

Mandible_length_min

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	Mandible_length_min
traitName	Mandible_length_min
trait_class	morphology
broaderTerm	https://ecologicaltraitdata.github.io/TraitDataList/#Mandible_length
valueType	numeric
traitUnit	mm
maxAllowedValue	999
minAllowedValue	0
factorLevels	NA
traitDescription	minimum documented length of a single mandible
Comments
example	0.01
relationSource	inheritsfrom
source	https://ecologicaltraitdata.github.io/TraitDataList/#Mandible_length

Tongue_length

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	Tongue_length
traitName	Tongue_length
trait_class	morphology
broaderTerm	https://ecologicaltraitdata.github.io/TraitDataList/#Nutrition_morphology
valueType	numeric
traitUnit	mm
maxAllowedValue	999
minAllowedValue	0
factorLevels	NA
traitDescription	length of the tounge
Comments
example	7
relationSource
source

Tongue_length_max

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	Tongue_length_max
traitName	Tongue_length_max
trait_class	morphology
broaderTerm	https://ecologicaltraitdata.github.io/TraitDataList/#Tongue_length
valueType	numeric
traitUnit	mm
maxAllowedValue	999
minAllowedValue	0
factorLevels	NA
traitDescription	maximum documented length of the tounge
Comments
example	8
relationSource	inheritsfrom
source	https://ecologicaltraitdata.github.io/TraitDataList/#Tongue_length

Tongue_length_min

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	Tongue_length_min
traitName	Tongue_length_min
trait_class	morphology
broaderTerm	https://ecologicaltraitdata.github.io/TraitDataList/#Tongue_length
valueType	numeric
traitUnit	mm
maxAllowedValue	999
minAllowedValue	0
factorLevels	NA
traitDescription	minimum documented length of the tounge
Comments
example	6
relationSource	inheritsfrom
source	https://ecologicaltraitdata.github.io/TraitDataList/#Tongue_length

Rostrum_length

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	Rostrum_length
traitName	Rostrum_length
trait_class	morphology
broaderTerm	https://ecologicaltraitdata.github.io/TraitDataList/#Nutrition_morphology
valueType	numeric
traitUnit	mm
maxAllowedValue	999
minAllowedValue	0
factorLevels	NA
traitDescription	length of the rostrum
Comments
example	5
relationSource
source

Rostrum_length_max

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	Rostrum_length_max
traitName	Rostrum_length_max
trait_class	morphology
broaderTerm	https://ecologicaltraitdata.github.io/TraitDataList/#Rostrum_length
valueType	numeric
traitUnit	mm
maxAllowedValue	999
minAllowedValue	0
factorLevels	NA
traitDescription	maximum documented length of the rostrum
Comments
example	6
relationSource	inheritsfrom
source	https://ecologicaltraitdata.github.io/TraitDataList/#Rostrum_length

Rostrum_length_min

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	Rostrum_length_min
traitName	Rostrum_length_min
trait_class	morphology
broaderTerm	https://ecologicaltraitdata.github.io/TraitDataList/#Rostrum_length
valueType	numeric
traitUnit	mm
maxAllowedValue	999
minAllowedValue	0
factorLevels	NA
traitDescription	minimum documented length of the rostrum
Comments
example	4
relationSource	inheritsfrom
source	https://ecologicaltraitdata.github.io/TraitDataList/#Rostrum_length

Rostrum_width

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	Rostrum_width
traitName	Rostrum_width
trait_class	morphology
broaderTerm	https://ecologicaltraitdata.github.io/TraitDataList/#Nutrition_morphology
valueType	numeric
traitUnit	mm
maxAllowedValue	999
minAllowedValue	0
factorLevels	NA
traitDescription	width of the rostrum
Comments
example	3
relationSource
source

Rostrum_width_max

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	Rostrum_width_max
traitName	Rostrum_width_max
trait_class	morphology
broaderTerm	https://ecologicaltraitdata.github.io/TraitDataList/#Rostrum_width
valueType	numeric
traitUnit	mm
maxAllowedValue	999
minAllowedValue	0
factorLevels	NA
traitDescription	maximum documented width of the rostrum
Comments
example	4
relationSource	inheritsfrom
source	https://ecologicaltraitdata.github.io/TraitDataList/#Rostrum_width

Rostrum_width_min

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	Rostrum_width_min
traitName	Rostrum_width_min
trait_class	morphology
broaderTerm	https://ecologicaltraitdata.github.io/TraitDataList/#Rostrum_width
valueType	numeric
traitUnit	mm
maxAllowedValue	999
minAllowedValue	0
factorLevels	NA
traitDescription	minimum documented width of the rostrum
Comments
example	2
relationSource	inheritsfrom
source	https://ecologicaltraitdata.github.io/TraitDataList/#Rostrum_width

Rostrum_ratio

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	Rostrum_ratio
traitName	Rostrum_ratio
trait_class	morphology
broaderTerm	https://ecologicaltraitdata.github.io/TraitDataList/#Nutrition_morphology
valueType	numeric
traitUnit	ratio
maxAllowedValue	999
minAllowedValue	0
factorLevels	NA
traitDescription	ratio between rostrum length and width
Comments
example	2
relationSource
source

Rostrum_ratio_max

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	Rostrum_ratio_max
traitName	Rostrum_ratio_max
trait_class	morphology
broaderTerm	https://ecologicaltraitdata.github.io/TraitDataList/#Rostrum_ratio
valueType	numeric
traitUnit	ratio
maxAllowedValue	999
minAllowedValue	0
factorLevels	NA
traitDescription	maximum documented ratio between rostrum length and width
Comments
example	2.5
relationSource	inheritsfrom
source	https://ecologicaltraitdata.github.io/TraitDataList/#Rostrum_ratio

Rostrum_ratio_min

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	Rostrum_ratio_min
traitName	Rostrum_ratio_min
trait_class	morphology
broaderTerm	https://ecologicaltraitdata.github.io/TraitDataList/#Rostrum_ratio
valueType	numeric
traitUnit	ratio
maxAllowedValue	999
minAllowedValue	0
factorLevels	NA
traitDescription	minimum documented ratio between rostrum length and width
Comments
example	1
relationSource	inheritsfrom
source	https://ecologicaltraitdata.github.io/TraitDataList/#Rostrum_ratio

Lipidic_storage

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	Lipidic_storage
traitName	Lipidic_storage
trait_class	morphology
broaderTerm	https://ecologicaltraitdata.github.io/TraitDataList/#Nutrition_morphology
valueType	character
traitUnit	NA
maxAllowedValue	NA
minAllowedValue	NA
factorLevels	NA
traitDescription	any morphological structure which serves a specialized function for nutrient storage within tissues
Comments
example	NA
relationSource	equals
source	http://t-sita.cesab.org/BETSI_vizInfo.jsp?trait=Lipidic_storage_tissues

Respiratory_organ_type

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	Respiratory_organ_type
traitName	Respiratory_organ_type
trait_class	morphology
broaderTerm	https://ecologicaltraitdata.github.io/TraitDataList/#morphology
valueType	factor
traitUnit	NA
maxAllowedValue	NA
minAllowedValue	NA
factorLevels	trachea/lung/dermis
traitDescription	type of respiratory organ
Comments	A collection of tissues joined in structural unit to the intake of oxygen and giving off of carbon dioxide which shares similar characteristics and forms a smaller division of a larger set
example	trachea
relationSource	equals
source	http://t-sita.cesab.org/BETSI_vizInfo.jsp?trait=Respiratory_organ_type

Sensory_morphology

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	Sensory_morphology
traitName	Sensory_morphology
trait_class	morphology
broaderTerm	https://ecologicaltraitdata.github.io/TraitDataList/#morphology
valueType	character
traitUnit	NA
maxAllowedValue	NA
minAllowedValue	NA
factorLevels	NA
traitDescription	any morphological structure which serves a specialized function for receiving sensory stimuli
Comments
example	NA
relationSource	equals
source	http://t-sita.cesab.org/BETSI_vizInfo.jsp?trait=Sensitivity

Antenna_length

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	Antenna_length
traitName	Antenna_length
trait_class	morphology
broaderTerm	https://ecologicaltraitdata.github.io/TraitDataList/#Sensory_morphology
valueType	numeric
traitUnit	cm
maxAllowedValue	999
minAllowedValue	0
factorLevels	NA
traitDescription	length of the complete antenna
Comments	usually measured by summing up the length of individual antenna segments
example	1.2
relationSource	equals
source	http://t-sita.cesab.org/BETSI_vizInfo.jsp?trait=Antenna_length

Antenna_length_max

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	Antenna_length_max
traitName	Antenna_length_max
trait_class	morphology
broaderTerm	https://ecologicaltraitdata.github.io/TraitDataList/#Antenna_length
valueType	numeric
traitUnit	cm
maxAllowedValue	999
minAllowedValue	0
factorLevels	NA
traitDescription	maximum documented length of the complete antenna
Comments	usually measured by summing up the length of individual antenna segments
example	1.2
relationSource	inheritsfrom
source	https://ecologicaltraitdata.github.io/TraitDataList/#Antenna_length

Antenna_length_min

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	Antenna_length_min
traitName	Antenna_length_min
trait_class	morphology
broaderTerm	https://ecologicaltraitdata.github.io/TraitDataList/#Antenna_length
valueType	numeric
traitUnit	cm
maxAllowedValue	999
minAllowedValue	0
factorLevels	NA
traitDescription	minimum documented length of the complete antenna
Comments	usually measured by summing up the length of individual antenna segments
example	1.2
relationSource	inheritsfrom
source	https://ecologicaltraitdata.github.io/TraitDataList/#Antenna_length

Antenna_segment1_length

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	Antenna_segment1_length
traitName	Antenna_segment1_length
trait_class	morphology
broaderTerm	https://ecologicaltraitdata.github.io/TraitDataList/#Antenna_length
valueType	numeric
traitUnit	cm
maxAllowedValue	999
minAllowedValue	0
factorLevels	NA
traitDescription	length of the first antenna segment
Comments	counted from the head
example	0.3
relationSource	inheritsfrom
source	https://ecologicaltraitdata.github.io/TraitDataList/#Antenna_length

Antenna_segment2_length

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	Antenna_segment2_length
traitName	Antenna_segment2_length
trait_class	morphology
broaderTerm	https://ecologicaltraitdata.github.io/TraitDataList/#Antenna_length
valueType	numeric
traitUnit	cm
maxAllowedValue	999
minAllowedValue	0
factorLevels	NA
traitDescription	length of the second antenna segment
Comments	counted from the head
example	0.25
relationSource	inheritsfrom
source	https://ecologicaltraitdata.github.io/TraitDataList/#Antenna_length

Antenna_segment3_length

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	Antenna_segment3_length
traitName	Antenna_segment3_length
trait_class	morphology
broaderTerm	https://ecologicaltraitdata.github.io/TraitDataList/#Antenna_length
valueType	numeric
traitUnit	cm
maxAllowedValue	999
minAllowedValue	0
factorLevels	NA
traitDescription	length of the third antenna segment
Comments	counted from the head
example	0.2
relationSource	inheritsfrom
source	https://ecologicaltraitdata.github.io/TraitDataList/#Antenna_length

Antenna_segment4_length

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	Antenna_segment4_length
traitName	Antenna_segment4_length
trait_class	morphology
broaderTerm	https://ecologicaltraitdata.github.io/TraitDataList/#Antenna_length
valueType	numeric
traitUnit	cm
maxAllowedValue	999
minAllowedValue	0
factorLevels	NA
traitDescription	length of the fourth antenna segment
Comments	counted from the head
example	0.15
relationSource	inheritsfrom
source	https://ecologicaltraitdata.github.io/TraitDataList/#Antenna_length

Antenna_segment5_length

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	Antenna_segment5_length
traitName	Antenna_segment5_length
trait_class	morphology
broaderTerm	https://ecologicaltraitdata.github.io/TraitDataList/#Antenna_length
valueType	numeric
traitUnit	cm
maxAllowedValue	999
minAllowedValue	0
factorLevels	NA
traitDescription	length of the fifth antenna segment
Comments	counted from the head
example	0.1
relationSource	inheritsfrom
source	https://ecologicaltraitdata.github.io/TraitDataList/#Antenna_length

Antenna_segment6_length

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	Antenna_segment6_length
traitName	Antenna_segment6_length
trait_class	morphology
broaderTerm	https://ecologicaltraitdata.github.io/TraitDataList/#Antenna_length
valueType	numeric
traitUnit	cm
maxAllowedValue	999
minAllowedValue	0
factorLevels	NA
traitDescription	length of the sixth antenna segment
Comments	counted from the head
example	0.05
relationSource	inheritsfrom
source	https://ecologicaltraitdata.github.io/TraitDataList/#Antenna_length

General_Body_pubescence

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	General_Body_pubescence
traitName	General_Body_pubescence
trait_class	morphology
broaderTerm	https://ecologicaltraitdata.github.io/TraitDataList/#Sensory_morphology
valueType	factor
traitUnit	NA
maxAllowedValue	NA
minAllowedValue	NA
factorLevels	none/sparse/dense/patchy
traitDescription	category of body pubescence
Comments
example	sparse
relationSource
source

Chemoreceptor

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	Chemoreceptor
traitName	Chemoreceptor
trait_class	morphology
broaderTerm	https://ecologicaltraitdata.github.io/TraitDataList/#Sensory_morphology
valueType	logical
traitUnit	NA
maxAllowedValue	NA
minAllowedValue	NA
factorLevels	NA
traitDescription	presence of chemoreceptors
Comments
example	TRUE
relationSource	equals
source	http://t-sita.cesab.org/BETSI_vizInfo.jsp?trait=Chemoreceptor

Thermoreceptor

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	Thermoreceptor
traitName	Thermoreceptor
trait_class	morphology
broaderTerm	https://ecologicaltraitdata.github.io/TraitDataList/#Sensory_morphology
valueType	logical
traitUnit	NA
maxAllowedValue	NA
minAllowedValue	NA
factorLevels	NA
traitDescription	presence of thermoreceptors
Comments
example	TRUE
relationSource	equals
source	http://t-sita.cesab.org/BETSI_vizInfo.jsp?trait=Thermoreceptor

Mechanoreceptor

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	Mechanoreceptor
traitName	Mechanoreceptor
trait_class	morphology
broaderTerm	https://ecologicaltraitdata.github.io/TraitDataList/#Sensory_morphology
valueType	logical
traitUnit	NA
maxAllowedValue	NA
minAllowedValue	NA
factorLevels	NA
traitDescription	presence of mechanoreceptors
Comments
example	TRUE
relationSource	equals
source	http://t-sita.cesab.org/BETSI_vizInfo.jsp?trait=Mechanoreceptor

Photoreceptor

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	Photoreceptor
traitName	Photoreceptor
trait_class	morphology
broaderTerm	https://ecologicaltraitdata.github.io/TraitDataList/#Sensory_morphology
valueType	logical
traitUnit	NA
maxAllowedValue	NA
minAllowedValue	NA
factorLevels	NA
traitDescription	presence of photoreceptors
Comments
example	TRUE
relationSource	equals
source	http://t-sita.cesab.org/BETSI_vizInfo.jsp?trait=Photoreceptor

Eye_number

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	Eye_number
traitName	Eye_number
trait_class	morphology
broaderTerm	https://ecologicaltraitdata.github.io/TraitDataList/#Photoreceptor
valueType	numeric
traitUnit	count
maxAllowedValue	999
minAllowedValue	0
factorLevels	NA
traitDescription	number of eyes
Comments
example	8
relationSource	equals
source	http://t-sita.cesab.org/BETSI_vizInfo.jsp?trait=Maximal-number-of%20-visual-organ

Eye_area

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	Eye_area
traitName	Eye_area
trait_class	morphology
broaderTerm	https://ecologicaltraitdata.github.io/TraitDataList/#Photoreceptor
valueType	numeric
traitUnit	mm²
maxAllowedValue	999
minAllowedValue	0
factorLevels	NA
traitDescription	total area comprised by all eyes
Comments	usually calculated from measures of eye width/length/diameter
example	1.6
relationSource	inheritsfrom
source	http://t-sita.cesab.org/BETSI_vizInfo.jsp?trait=Eye_diameter

Eye_width

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	Eye_width
traitName	Eye_width
trait_class	morphology
broaderTerm	https://ecologicaltraitdata.github.io/TraitDataList/#Photoreceptor
valueType	numeric
traitUnit	mm
maxAllowedValue	999
minAllowedValue	0
factorLevels	NA
traitDescription	width of a single eye
Comments
example	2
relationSource	inheritsfrom
source	http://t-sita.cesab.org/BETSI_vizInfo.jsp?trait=Eye_diameter

Eye_width_max

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	Eye_width_max
traitName	Eye_width_max
trait_class	morphology
broaderTerm	https://ecologicaltraitdata.github.io/TraitDataList/#Eye_width
valueType	numeric
traitUnit	mm
maxAllowedValue	999
minAllowedValue	0
factorLevels	NA
traitDescription	maximum documented width of a single eye
Comments
example	2.5
relationSource	inheritsfrom
source	https://ecologicaltraitdata.github.io/TraitDataList/#Eye_width

Eye_width_min

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	Eye_width_min
traitName	Eye_width_min
trait_class	morphology
broaderTerm	https://ecologicaltraitdata.github.io/TraitDataList/#Eye_width
valueType	numeric
traitUnit	mm
maxAllowedValue	999
minAllowedValue	0
factorLevels	NA
traitDescription	minimum documented width of a single eye
Comments
example	1.8
relationSource	inheritsfrom
source	https://ecologicaltraitdata.github.io/TraitDataList/#Eye_width

Eye_diameter

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	Eye_diameter
traitName	Eye_diameter
trait_class	morphology
broaderTerm	https://ecologicaltraitdata.github.io/TraitDataList/#Photoreceptor
valueType	numeric
traitUnit	mm
maxAllowedValue	999
minAllowedValue	0
factorLevels	NA
traitDescription	diameter of a single eye
Comments
example	2
relationSource	equals
source	http://t-sita.cesab.org/BETSI_vizInfo.jsp?trait=Eye_diameter

Eye_diameter_max

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	Eye_diameter_max
traitName	Eye_diameter_max
trait_class	morphology
broaderTerm	https://ecologicaltraitdata.github.io/TraitDataList/#Eye_diameter
valueType	numeric
traitUnit	mm
maxAllowedValue	999
minAllowedValue	0
factorLevels	NA
traitDescription	maximum documented diameter of a single eye
Comments
example	2.5
relationSource	inheritsfrom
source	https://ecologicaltraitdata.github.io/TraitDataList/#Eye_diameter

Eye_diameter_min

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	Eye_diameter_min
traitName	Eye_diameter_min
trait_class	morphology
broaderTerm	https://ecologicaltraitdata.github.io/TraitDataList/#Eye_diameter
valueType	numeric
traitUnit	mm
maxAllowedValue	999
minAllowedValue	0
factorLevels	NA
traitDescription	minimum documented diameter of a single eye
Comments
example	1.8
relationSource	inheritsfrom
source	https://ecologicaltraitdata.github.io/TraitDataList/#Eye_diameter

Locomotion_morphology

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	Locomotion_morphology
traitName	Locomotion_morphology
trait_class	morphology
broaderTerm	https://ecologicaltraitdata.github.io/TraitDataList/#morphology
valueType	character
traitUnit	NA
maxAllowedValue	NA
minAllowedValue	NA
factorLevels	NA
traitDescription	any morphological structure which serves a specialized function for locomotion
Comments
example	NA
relationSource	equals
source	http://t-sita.cesab.org/BETSI_vizInfo.jsp?trait=Locomotion

Leg_length

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	Leg_length
traitName	Leg_length
trait_class	morphology
broaderTerm	https://ecologicaltraitdata.github.io/TraitDataList/#Locomotion_morphology
valueType	numeric
traitUnit	mm
maxAllowedValue	999
minAllowedValue	0
factorLevels	NA
traitDescription	total length of the leg
Comments
example	25
relationSource	equals
source	http://t-sita.cesab.org/BETSI_vizInfo.jsp?trait=leg_length

Leg_length_max

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	Leg_length_max
traitName	Leg_length_max
trait_class	morphology
broaderTerm	https://ecologicaltraitdata.github.io/TraitDataList/#Leg_length
valueType	numeric
traitUnit	mm
maxAllowedValue	999
minAllowedValue	0
factorLevels	NA
traitDescription	maximum documented total length of the leg
Comments
example	30
relationSource	inheritsfrom
source	https://ecologicaltraitdata.github.io/TraitDataList/#Leg_length

Leg_length_min

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	Leg_length_min
traitName	Leg_length_min
trait_class	morphology
broaderTerm	https://ecologicaltraitdata.github.io/TraitDataList/#Leg_length
valueType	numeric
traitUnit	mm
maxAllowedValue	999
minAllowedValue	0
factorLevels	NA
traitDescription	minimum documented total length of the leg
Comments
example	20
relationSource	inheritsfrom
source	https://ecologicaltraitdata.github.io/TraitDataList/#Leg_length

Hind_leg_length

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	Hind_leg_length
traitName	Hind_leg_length
trait_class	morphology
broaderTerm	https://ecologicaltraitdata.github.io/TraitDataList/#Leg_length
valueType	numeric
traitUnit	mm
maxAllowedValue	999
minAllowedValue	0
factorLevels	NA
traitDescription	total length of the hind leg
Comments
example	25
relationSource	refines
source	http://t-sita.cesab.org/BETSI_vizInfo.jsp?trait=leg_length

Hind_leg_length_max

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	Hind_leg_length_max
traitName	Hind_leg_length_max
trait_class	morphology
broaderTerm	https://ecologicaltraitdata.github.io/TraitDataList/#Hind_leg_length
valueType	numeric
traitUnit	mm
maxAllowedValue	999
minAllowedValue	0
factorLevels	NA
traitDescription	maximum documented total length of the hind leg
Comments
example	30
relationSource	inheritsfrom
source	https://ecologicaltraitdata.github.io/TraitDataList/#Hind_leg_length

Hind_leg_length_min

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	Hind_leg_length_min
traitName	Hind_leg_length_min
trait_class	morphology
broaderTerm	https://ecologicaltraitdata.github.io/TraitDataList/#Hind_leg_length
valueType	numeric
traitUnit	mm
maxAllowedValue	999
minAllowedValue	0
factorLevels	NA
traitDescription	minimum documented total length of the hind leg
Comments
example	20
relationSource	inheritsfrom
source	https://ecologicaltraitdata.github.io/TraitDataList/#Hind_leg_length

Mid_leg_length

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	Mid_leg_length
traitName	Mid_leg_length
trait_class	morphology
broaderTerm	https://ecologicaltraitdata.github.io/TraitDataList/#Leg_length
valueType	numeric
traitUnit	mm
maxAllowedValue	999
minAllowedValue	0
factorLevels	NA
traitDescription	total length of the mid leg
Comments
example	25
relationSource	refines
source	http://t-sita.cesab.org/BETSI_vizInfo.jsp?trait=leg_length

Mid_leg_length_max

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	Mid_leg_length_max
traitName	Mid_leg_length_max
trait_class	morphology
broaderTerm	https://ecologicaltraitdata.github.io/TraitDataList/#Mid_leg_length
valueType	numeric
traitUnit	mm
maxAllowedValue	999
minAllowedValue	0
factorLevels	NA
traitDescription	maximum documented total length of the mid leg
Comments
example	30
relationSource	inheritsfrom
source	https://ecologicaltraitdata.github.io/TraitDataList/#Mid_leg_length

Mid_leg_length_min

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	Mid_leg_length_min
traitName	Mid_leg_length_min
trait_class	morphology
broaderTerm	https://ecologicaltraitdata.github.io/TraitDataList/#Mid_leg_length
valueType	numeric
traitUnit	mm
maxAllowedValue	999
minAllowedValue	0
factorLevels	NA
traitDescription	minimum documented total length of the mid leg
Comments
example	20
relationSource	inheritsfrom
source	https://ecologicaltraitdata.github.io/TraitDataList/#Mid_leg_length

Front_leg_length

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	Front_leg_length
traitName	Front_leg_length
trait_class	morphology
broaderTerm	https://ecologicaltraitdata.github.io/TraitDataList/#Leg_length
valueType	numeric
traitUnit	mm
maxAllowedValue	999
minAllowedValue	0
factorLevels	NA
traitDescription	total length of the front leg
Comments
example	25
relationSource	refines
source	http://t-sita.cesab.org/BETSI_vizInfo.jsp?trait=leg_length

Front_leg_length_max

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	Front_leg_length_max
traitName	Front_leg_length_max
trait_class	morphology
broaderTerm	https://ecologicaltraitdata.github.io/TraitDataList/#Front_leg_length
valueType	numeric
traitUnit	mm
maxAllowedValue	999
minAllowedValue	0
factorLevels	NA
traitDescription	maximum documented total length of the front leg
Comments
example	30
relationSource	inheritsfrom
source	https://ecologicaltraitdata.github.io/TraitDataList/#Front_leg_length

Front_leg_length_min

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	Front_leg_length_min
traitName	Front_leg_length_min
trait_class	morphology
broaderTerm	https://ecologicaltraitdata.github.io/TraitDataList/#Front_leg_length
valueType	numeric
traitUnit	mm
maxAllowedValue	999
minAllowedValue	0
factorLevels	NA
traitDescription	minimum documented total length of the front leg
Comments
example	20
relationSource	inheritsfrom
source	https://ecologicaltraitdata.github.io/TraitDataList/#Front_leg_length

Femur_length

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	Femur_length
traitName	Femur_length
trait_class	morphology
broaderTerm	https://ecologicaltraitdata.github.io/TraitDataList/#Leg_length
valueType	numeric
traitUnit	mm
maxAllowedValue	999
minAllowedValue	0
factorLevels	NA
traitDescription	length of femur
Comments
example	15
relationSource	equals
source	http://t-sita.cesab.org/BETSI_vizInfo.jsp?trait=Femur_length

Femur_length_max

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	Femur_length_max
traitName	Femur_length_max
trait_class	morphology
broaderTerm	https://ecologicaltraitdata.github.io/TraitDataList/#Femur_length
valueType	numeric
traitUnit	mm
maxAllowedValue	999
minAllowedValue	0
factorLevels	NA
traitDescription	maximum documented length of femur
Comments
example	17
relationSource	inheritsfrom
source	https://ecologicaltraitdata.github.io/TraitDataList/#Femur_length

Femur_length_min

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	Femur_length_min
traitName	Femur_length_min
trait_class	morphology
broaderTerm	https://ecologicaltraitdata.github.io/TraitDataList/#Femur_length
valueType	numeric
traitUnit	mm
maxAllowedValue	999
minAllowedValue	0
factorLevels	NA
traitDescription	minimum documented length of femur
Comments
example	13
relationSource	inheritsfrom
source	https://ecologicaltraitdata.github.io/TraitDataList/#Femur_length

Front_femur_length

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	Front_femur_length
traitName	Front_femur_length
trait_class	morphology
broaderTerm	https://ecologicaltraitdata.github.io/TraitDataList/#Femur_length
valueType	numeric
traitUnit	mm
maxAllowedValue	999
minAllowedValue	0
factorLevels	NA
traitDescription	length of front femur
Comments
example	15
relationSource	refines
source	http://t-sita.cesab.org/BETSI_vizInfo.jsp?trait=Femur_length

Front_femur_length_max

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	Front_femur_length_max
traitName	Front_femur_length_max
trait_class	morphology
broaderTerm	https://ecologicaltraitdata.github.io/TraitDataList/#Front_femur_length
valueType	numeric
traitUnit	mm
maxAllowedValue	999
minAllowedValue	0
factorLevels	NA
traitDescription	maximum documented length of front femur
Comments
example	17
relationSource	inheritsfrom
source	https://ecologicaltraitdata.github.io/TraitDataList/#Front_femur_length

Front_femur_length_min

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	Front_femur_length_min
traitName	Front_femur_length_min
trait_class	morphology
broaderTerm	https://ecologicaltraitdata.github.io/TraitDataList/#Front_femur_length
valueType	numeric
traitUnit	mm
maxAllowedValue	999
minAllowedValue	0
factorLevels	NA
traitDescription	minimum documented length of front femur
Comments
example	13
relationSource	inheritsfrom
source	https://ecologicaltraitdata.github.io/TraitDataList/#Front_femur_length

Mid_femur_length

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	Mid_femur_length
traitName	Mid_femur_length
trait_class	morphology
broaderTerm	https://ecologicaltraitdata.github.io/TraitDataList/#Femur_length
valueType	numeric
traitUnit	mm
maxAllowedValue	999
minAllowedValue	0
factorLevels	NA
traitDescription	length of mid femur
Comments
example	15
relationSource	refines
source	http://t-sita.cesab.org/BETSI_vizInfo.jsp?trait=Femur_length

Mid_femur_length_max

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	Mid_femur_length_max
traitName	Mid_femur_length_max
trait_class	morphology
broaderTerm	https://ecologicaltraitdata.github.io/TraitDataList/#Mid_femur_length
valueType	numeric
traitUnit	mm
maxAllowedValue	999
minAllowedValue	0
factorLevels	NA
traitDescription	maximum documented length of mid femur
Comments
example	17
relationSource	inheritsfrom
source	https://ecologicaltraitdata.github.io/TraitDataList/#Mid_femur_length

Mid_femur_length_min

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	Mid_femur_length_min
traitName	Mid_femur_length_min
trait_class	morphology
broaderTerm	https://ecologicaltraitdata.github.io/TraitDataList/#Mid_femur_length
valueType	numeric
traitUnit	mm
maxAllowedValue	999
minAllowedValue	0
factorLevels	NA
traitDescription	minimum documented length of mid femur
Comments
example	13
relationSource	inheritsfrom
source	https://ecologicaltraitdata.github.io/TraitDataList/#Mid_femur_length

Hind_femur_length

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	Hind_femur_length
traitName	Hind_femur_length
trait_class	morphology
broaderTerm	https://ecologicaltraitdata.github.io/TraitDataList/#Femur_length
valueType	numeric
traitUnit	mm
maxAllowedValue	999
minAllowedValue	0
factorLevels	NA
traitDescription	length of hind femur
Comments
example	15
relationSource	refines
source	http://t-sita.cesab.org/BETSI_vizInfo.jsp?trait=Femur_length

Hind_femur_length_max

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	Hind_femur_length_max
traitName	Hind_femur_length_max
trait_class	morphology
broaderTerm	https://ecologicaltraitdata.github.io/TraitDataList/#Hind_femur_length
valueType	numeric
traitUnit	mm
maxAllowedValue	999
minAllowedValue	0
factorLevels	NA
traitDescription	maximum documented length of hind femur
Comments
example	17
relationSource	inheritsfrom
source	https://ecologicaltraitdata.github.io/TraitDataList/#Hind_femur_length

Hind_femur_length_min

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	Hind_femur_length_min
traitName	Hind_femur_length_min
trait_class	morphology
broaderTerm	https://ecologicaltraitdata.github.io/TraitDataList/#Hind_femur_length
valueType	numeric
traitUnit	mm
maxAllowedValue	999
minAllowedValue	0
factorLevels	NA
traitDescription	minimum documented length of hind femur
Comments
example	13
relationSource	inheritsfrom
source	https://ecologicaltraitdata.github.io/TraitDataList/#Hind_femur_length

Tibia_length

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	Tibia_length
traitName	Tibia_length
trait_class	morphology
broaderTerm	https://ecologicaltraitdata.github.io/TraitDataList/#Leg_length
valueType	numeric
traitUnit	mm
maxAllowedValue	999
minAllowedValue	0
factorLevels	NA
traitDescription	length of Tibia
Comments
example	15
relationSource	equals
source	http://t-sita.cesab.org/BETSI_vizInfo.jsp?trait=Tibia_length

Tibia_length_max

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	Tibia_length_max
traitName	Tibia_length_max
trait_class	morphology
broaderTerm	https://ecologicaltraitdata.github.io/TraitDataList/#Tibia_length
valueType	numeric
traitUnit	mm
maxAllowedValue	999
minAllowedValue	0
factorLevels	NA
traitDescription	maximum documented length of Tibia
Comments
example	17
relationSource	inheritsfrom
source	https://ecologicaltraitdata.github.io/TraitDataList/#Tibia_length

Tibia_length_min

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	Tibia_length_min
traitName	Tibia_length_min
trait_class	morphology
broaderTerm	https://ecologicaltraitdata.github.io/TraitDataList/#Tibia_length
valueType	numeric
traitUnit	mm
maxAllowedValue	999
minAllowedValue	0
factorLevels	NA
traitDescription	minimum documented length of Tibia
Comments
example	13
relationSource	inheritsfrom
source	https://ecologicaltraitdata.github.io/TraitDataList/#Tibia_length

Front_tibia_length

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	Front_tibia_length
traitName	Front_tibia_length
trait_class	morphology
broaderTerm	https://ecologicaltraitdata.github.io/TraitDataList/#Tibia_length
valueType	numeric
traitUnit	mm
maxAllowedValue	999
minAllowedValue	0
factorLevels	NA
traitDescription	length of front tibia
Comments
example	15
relationSource	refines
source	http://t-sita.cesab.org/BETSI_vizInfo.jsp?trait=Tibia_length

Front_tibia_length_max

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	Front_tibia_length_max
traitName	Front_tibia_length_max
trait_class	morphology
broaderTerm	https://ecologicaltraitdata.github.io/TraitDataList/#Front_tibia_length
valueType	numeric
traitUnit	mm
maxAllowedValue	999
minAllowedValue	0
factorLevels	NA
traitDescription	maximum documented length of front tibia
Comments
example	17
relationSource	inheritsfrom
source	https://ecologicaltraitdata.github.io/TraitDataList/#Front_tibia_length

Front_tibia_length_min

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	Front_tibia_length_min
traitName	Front_tibia_length_min
trait_class	morphology
broaderTerm	https://ecologicaltraitdata.github.io/TraitDataList/#Front_tibia_length
valueType	numeric
traitUnit	mm
maxAllowedValue	999
minAllowedValue	0
factorLevels	NA
traitDescription	minimum documented length of front tibia
Comments
example	13
relationSource	inheritsfrom
source	https://ecologicaltraitdata.github.io/TraitDataList/#Front_tibia_length

Mid_Tibia_length

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	Mid_Tibia_length
traitName	Mid_Tibia_length
trait_class	morphology
broaderTerm	https://ecologicaltraitdata.github.io/TraitDataList/#Tibia_length
valueType	numeric
traitUnit	mm
maxAllowedValue	999
minAllowedValue	0
factorLevels	NA
traitDescription	length of mid tibia
Comments
example	15
relationSource	refines
source	http://t-sita.cesab.org/BETSI_vizInfo.jsp?trait=Tibia_length

Mid_Tibia_length_max

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	Mid_Tibia_length_max
traitName	Mid_Tibia_length_max
trait_class	morphology
broaderTerm	https://ecologicaltraitdata.github.io/TraitDataList/#Mid_Tibia_length
valueType	numeric
traitUnit	mm
maxAllowedValue	999
minAllowedValue	0
factorLevels	NA
traitDescription	maximum documented length of mid tibia
Comments
example	17
relationSource	inheritsfrom
source	https://ecologicaltraitdata.github.io/TraitDataList/#Mid_Tibia_length

Mid_Tibia_length_min

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	Mid_Tibia_length_min
traitName	Mid_Tibia_length_min
trait_class	morphology
broaderTerm	https://ecologicaltraitdata.github.io/TraitDataList/#Mid_Tibia_length
valueType	numeric
traitUnit	mm
maxAllowedValue	999
minAllowedValue	0
factorLevels	NA
traitDescription	minimum documented length of mid tibia
Comments
example	13
relationSource	inheritsfrom
source	https://ecologicaltraitdata.github.io/TraitDataList/#Mid_Tibia_length

Hind_tibia_length

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	Hind_tibia_length
traitName	Hind_tibia_length
trait_class	morphology
broaderTerm	https://ecologicaltraitdata.github.io/TraitDataList/#Tibia_length
valueType	numeric
traitUnit	mm
maxAllowedValue	999
minAllowedValue	0
factorLevels	NA
traitDescription	length of hind tibia
Comments
example	15
relationSource	refines
source	http://t-sita.cesab.org/BETSI_vizInfo.jsp?trait=Tibia_length

Hind_tibia_length_max

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	Hind_tibia_length_max
traitName	Hind_tibia_length_max
trait_class	morphology
broaderTerm	https://ecologicaltraitdata.github.io/TraitDataList/#Hind_tibia_length
valueType	numeric
traitUnit	mm
maxAllowedValue	999
minAllowedValue	0
factorLevels	NA
traitDescription	maximum documented length of hind tibia
Comments
example	17
relationSource	inheritsfrom
source	https://ecologicaltraitdata.github.io/TraitDataList/#Hind_tibia_length

Hind_tibia_length_min

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	Hind_tibia_length_min
traitName	Hind_tibia_length_min
trait_class	morphology
broaderTerm	https://ecologicaltraitdata.github.io/TraitDataList/#Hind_tibia_length
valueType	numeric
traitUnit	mm
maxAllowedValue	999
minAllowedValue	0
factorLevels	NA
traitDescription	minimum documented length of hind tibia
Comments
example	13
relationSource	inheritsfrom
source	https://ecologicaltraitdata.github.io/TraitDataList/#Hind_tibia_length

Trochanter_length

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	Trochanter_length
traitName	Trochanter_length
trait_class	morphology
broaderTerm	https://ecologicaltraitdata.github.io/TraitDataList/#Leg_length
valueType	numeric
traitUnit	mm
maxAllowedValue	999
minAllowedValue	0
factorLevels	NA
traitDescription	length of trochanter
Comments
example	15
relationSource	equals
source	http://t-sita.cesab.org/BETSI_vizInfo.jsp?trait=Trochanter_length

Trochanter_length_max

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	Trochanter_length_max
traitName	Trochanter_length_max
trait_class	morphology
broaderTerm	https://ecologicaltraitdata.github.io/TraitDataList/#Trochanter_length
valueType	numeric
traitUnit	mm
maxAllowedValue	999
minAllowedValue	0
factorLevels	NA
traitDescription	maximum documented length of trochanter
Comments
example	17
relationSource	inheritsfrom
source	https://ecologicaltraitdata.github.io/TraitDataList/#Trochanter_length

Trochanter_length_min

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	Trochanter_length_min
traitName	Trochanter_length_min
trait_class	morphology
broaderTerm	https://ecologicaltraitdata.github.io/TraitDataList/#Trochanter_length
valueType	numeric
traitUnit	mm
maxAllowedValue	999
minAllowedValue	0
factorLevels	NA
traitDescription	minimum documented length of trochanter
Comments
example	13
relationSource	inheritsfrom
source	https://ecologicaltraitdata.github.io/TraitDataList/#Trochanter_length

Front_trochanter_length

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	Front_trochanter_length
traitName	Front_trochanter_length
trait_class	morphology
broaderTerm	https://ecologicaltraitdata.github.io/TraitDataList/#Trochanter_length
valueType	numeric
traitUnit	mm
maxAllowedValue	999
minAllowedValue	0
factorLevels	NA
traitDescription	length of front trochanter
Comments
example	15
relationSource	refines
source	http://t-sita.cesab.org/BETSI_vizInfo.jsp?trait=Trochanter_length

Front_trochanter_length_max

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	Front_trochanter_length_max
traitName	Front_trochanter_length_max
trait_class	morphology
broaderTerm	https://ecologicaltraitdata.github.io/TraitDataList/#Front_trochanter_length
valueType	numeric
traitUnit	mm
maxAllowedValue	999
minAllowedValue	0
factorLevels	NA
traitDescription	maximum documented length of front trochanter
Comments
example	17
relationSource	inheritsfrom
source	https://ecologicaltraitdata.github.io/TraitDataList/#Front_trochanter_length

Front_trochanter_length_min

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	Front_trochanter_length_min
traitName	Front_trochanter_length_min
trait_class	morphology
broaderTerm	https://ecologicaltraitdata.github.io/TraitDataList/#Front_trochanter_length
valueType	numeric
traitUnit	mm
maxAllowedValue	999
minAllowedValue	0
factorLevels	NA
traitDescription	minimum documented length of front trochanter
Comments
example	13
relationSource	inheritsfrom
source	https://ecologicaltraitdata.github.io/TraitDataList/#Front_trochanter_length

Mid_trochanter_length

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	Mid_trochanter_length
traitName	Mid_trochanter_length
trait_class	morphology
broaderTerm	https://ecologicaltraitdata.github.io/TraitDataList/#Trochanter_length
valueType	numeric
traitUnit	mm
maxAllowedValue	999
minAllowedValue	0
factorLevels	NA
traitDescription	length of mid trochanter
Comments
example	15
relationSource	refines
source	http://t-sita.cesab.org/BETSI_vizInfo.jsp?trait=Trochanter_length

Mid_trochanter_length_max

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	Mid_trochanter_length_max
traitName	Mid_trochanter_length_max
trait_class	morphology
broaderTerm	https://ecologicaltraitdata.github.io/TraitDataList/#Mid_trochanter_length
valueType	numeric
traitUnit	mm
maxAllowedValue	999
minAllowedValue	0
factorLevels	NA
traitDescription	maximum documented length of mid trochanter
Comments
example	17
relationSource	inheritsfrom
source	https://ecologicaltraitdata.github.io/TraitDataList/#Mid_trochanter_length

Mid_trochanter_length_min

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	Mid_trochanter_length_min
traitName	Mid_trochanter_length_min
trait_class	morphology
broaderTerm	https://ecologicaltraitdata.github.io/TraitDataList/#Mid_trochanter_length
valueType	numeric
traitUnit	mm
maxAllowedValue	999
minAllowedValue	0
factorLevels	NA
traitDescription	minimum documented length of mid trochanter
Comments
example	13
relationSource	inheritsfrom
source	https://ecologicaltraitdata.github.io/TraitDataList/#Mid_trochanter_length

Hind_trochanter_length

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	Hind_trochanter_length
traitName	Hind_trochanter_length
trait_class	morphology
broaderTerm	https://ecologicaltraitdata.github.io/TraitDataList/#Trochanter_length
valueType	numeric
traitUnit	mm
maxAllowedValue	999
minAllowedValue	0
factorLevels	NA
traitDescription	length of hind trochanter
Comments
example	15
relationSource	refines
source	http://t-sita.cesab.org/BETSI_vizInfo.jsp?trait=Trochanter_length

Hind_trochanter_length_max

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	Hind_trochanter_length_max
traitName	Hind_trochanter_length_max
trait_class	morphology
broaderTerm	https://ecologicaltraitdata.github.io/TraitDataList/#Hind_trochanter_length
valueType	numeric
traitUnit	mm
maxAllowedValue	999
minAllowedValue	0
factorLevels	NA
traitDescription	maximum documented length of hind trochanter
Comments
example	17
relationSource	inheritsfrom
source	https://ecologicaltraitdata.github.io/TraitDataList/#Hind_trochanter_length

Hind_trochanter_length_min

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	Hind_trochanter_length_min
traitName	Hind_trochanter_length_min
trait_class	morphology
broaderTerm	https://ecologicaltraitdata.github.io/TraitDataList/#Hind_trochanter_length
valueType	numeric
traitUnit	mm
maxAllowedValue	999
minAllowedValue	0
factorLevels	NA
traitDescription	minimum documented length of hind trochanter
Comments
example	13
relationSource	inheritsfrom
source	https://ecologicaltraitdata.github.io/TraitDataList/#Hind_trochanter_length

Femur_width

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	Femur_width
traitName	Femur_width
trait_class	morphology
broaderTerm	https://ecologicaltraitdata.github.io/TraitDataList/#Locomotion_morphology
valueType	numeric
traitUnit	mm
maxAllowedValue	999
minAllowedValue	0
factorLevels	NA
traitDescription	width of the femur
Comments
example	3
relationSource	equals
source	http://t-sita.cesab.org/BETSI_vizInfo.jsp?trait=Femur_width

Femur_width_max

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	Femur_width_max
traitName	Femur_width_max
trait_class	morphology
broaderTerm	https://ecologicaltraitdata.github.io/TraitDataList/#Femur_width
valueType	numeric
traitUnit	mm
maxAllowedValue	999
minAllowedValue	0
factorLevels	NA
traitDescription	maximum documented width of the femur
Comments
example	4
relationSource	inheritsfrom
source	https://ecologicaltraitdata.github.io/TraitDataList/#Femur_width

Femur_width_min

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	Femur_width_min
traitName	Femur_width_min
trait_class	morphology
broaderTerm	https://ecologicaltraitdata.github.io/TraitDataList/#Femur_width
valueType	numeric
traitUnit	mm
maxAllowedValue	999
minAllowedValue	0
factorLevels	NA
traitDescription	minimum documented width of the femur
Comments
example	2
relationSource	inheritsfrom
source	https://ecologicaltraitdata.github.io/TraitDataList/#Femur_width

Front_femur_width

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	Front_femur_width
traitName	Front_femur_width
trait_class	morphology
broaderTerm	https://ecologicaltraitdata.github.io/TraitDataList/#Femur_width
valueType	numeric
traitUnit	mm
maxAllowedValue	999
minAllowedValue	0
factorLevels	NA
traitDescription	width of the front femur
Comments
example	3
relationSource	refines
source	http://t-sita.cesab.org/BETSI_vizInfo.jsp?trait=Femur_width

Front_femur_width_max

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	Front_femur_width_max
traitName	Front_femur_width_max
trait_class	morphology
broaderTerm	https://ecologicaltraitdata.github.io/TraitDataList/#Front_femur_width
valueType	numeric
traitUnit	mm
maxAllowedValue	999
minAllowedValue	0
factorLevels	NA
traitDescription	maximum documented width of the front femur
Comments
example	4
relationSource	inheritsfrom
source	https://ecologicaltraitdata.github.io/TraitDataList/#Front_femur_width

Front_femur_width_min

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	Front_femur_width_min
traitName	Front_femur_width_min
trait_class	morphology
broaderTerm	https://ecologicaltraitdata.github.io/TraitDataList/#Front_femur_width
valueType	numeric
traitUnit	mm
maxAllowedValue	999
minAllowedValue	0
factorLevels	NA
traitDescription	minimum documented width of the front femur
Comments
example	2
relationSource	inheritsfrom
source	https://ecologicaltraitdata.github.io/TraitDataList/#Front_femur_width

Mid_femur_width

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	Mid_femur_width
traitName	Mid_femur_width
trait_class	morphology
broaderTerm	https://ecologicaltraitdata.github.io/TraitDataList/#Femur_width
valueType	numeric
traitUnit	mm
maxAllowedValue	999
minAllowedValue	0
factorLevels	NA
traitDescription	width of the mid femur
Comments
example	3
relationSource	refines
source	http://t-sita.cesab.org/BETSI_vizInfo.jsp?trait=Femur_width

Mid_femur_width_max

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	Mid_femur_width_max
traitName	Mid_femur_width_max
trait_class	morphology
broaderTerm	https://ecologicaltraitdata.github.io/TraitDataList/#Mid_femur_width
valueType	numeric
traitUnit	mm
maxAllowedValue	999
minAllowedValue	0
factorLevels	NA
traitDescription	maximum documented width of the mid femur
Comments
example	4
relationSource	inheritsfrom
source	https://ecologicaltraitdata.github.io/TraitDataList/#Mid_femur_width

Mid_femur_width_min

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	Mid_femur_width_min
traitName	Mid_femur_width_min
trait_class	morphology
broaderTerm	https://ecologicaltraitdata.github.io/TraitDataList/#Mid_femur_width
valueType	numeric
traitUnit	mm
maxAllowedValue	999
minAllowedValue	0
factorLevels	NA
traitDescription	minimum documented width of the mid femur
Comments
example	2
relationSource	inheritsfrom
source	https://ecologicaltraitdata.github.io/TraitDataList/#Mid_femur_width

Hind_femur_width

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	Hind_femur_width
traitName	Hind_femur_width
trait_class	morphology
broaderTerm	https://ecologicaltraitdata.github.io/TraitDataList/#Femur_width
valueType	numeric
traitUnit	mm
maxAllowedValue	999
minAllowedValue	0
factorLevels	NA
traitDescription	width of the hind femur
Comments
example	3
relationSource	refines
source	http://t-sita.cesab.org/BETSI_vizInfo.jsp?trait=Femur_width

Hind_femur_width_max

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	Hind_femur_width_max
traitName	Hind_femur_width_max
trait_class	morphology
broaderTerm	https://ecologicaltraitdata.github.io/TraitDataList/#Hind_femur_width
valueType	numeric
traitUnit	mm
maxAllowedValue	999
minAllowedValue	0
factorLevels	NA
traitDescription	maximum documented width of the hind femur
Comments
example	4
relationSource	inheritsfrom
source	https://ecologicaltraitdata.github.io/TraitDataList/#Hind_femur_width

Hind_femur_width_min

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	Hind_femur_width_min
traitName	Hind_femur_width_min
trait_class	morphology
broaderTerm	https://ecologicaltraitdata.github.io/TraitDataList/#Hind_femur_width
valueType	numeric
traitUnit	mm
maxAllowedValue	999
minAllowedValue	0
factorLevels	NA
traitDescription	minimum documented width of the hind femur
Comments
example	2
relationSource	inheritsfrom
source	https://ecologicaltraitdata.github.io/TraitDataList/#Hind_femur_width

Wing_appearance

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	Wing_appearance
traitName	Wing_appearance
trait_class	morphology
broaderTerm	https://ecologicaltraitdata.github.io/TraitDataList/#Locomotion_morphology
valueType	factor
traitUnit	NA
maxAllowedValue	NA
minAllowedValue	NA
factorLevels	apter/hypopter/submacropter/macropter/brachypter
traitDescription	type of membranous wings
Comments
example	apter
relationSource	refines
source	http://t-sita.cesab.org/BETSI_vizInfo.jsp?trait=Wing_development

Wing_size_relation

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	Wing_size_relation
traitName	Wing_size_relation
trait_class	morphology
broaderTerm	https://ecologicaltraitdata.github.io/TraitDataList/#Wing_appearance
valueType	factor
traitUnit	NA
maxAllowedValue	NA
minAllowedValue	NA
factorLevels	eupter/parapter
traitDescription	Parapter: front wings smaller than hind wings / Eupter: front wings equal with hind wings
Comments
example	eupter
relationSource
source

Front_wing_width

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	Front_wing_width
traitName	Front_wing_width
trait_class	morphology
broaderTerm	https://ecologicaltraitdata.github.io/TraitDataList/#Wing_size_relation
valueType	numeric
traitUnit	cm
maxAllowedValue	999
minAllowedValue	0
factorLevels	NA
traitDescription	second longest dimension of the front wing
Comments
example	2
relationSource	refines
source	http://t-sita.cesab.org/BETSI_vizInfo.jsp?trait=Wing

Front_wing_width_max

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	Front_wing_width_max
traitName	Front_wing_width_max
trait_class	morphology
broaderTerm	https://ecologicaltraitdata.github.io/TraitDataList/#Front_wing_width
valueType	numeric
traitUnit	cm
maxAllowedValue	999
minAllowedValue	0
factorLevels	NA
traitDescription	maximum documented second longest dimension of the front wing
Comments
example	2.5
relationSource	inheritsfrom
source	https://ecologicaltraitdata.github.io/TraitDataList/#Front_wing_width

Front_wing_width_min

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	Front_wing_width_min
traitName	Front_wing_width_min
trait_class	morphology
broaderTerm	https://ecologicaltraitdata.github.io/TraitDataList/#Front_wing_width
valueType	numeric
traitUnit	cm
maxAllowedValue	999
minAllowedValue	0
factorLevels	NA
traitDescription	minimum documented second longest dimension of the front wing
Comments
example	1.5
relationSource	inheritsfrom
source	https://ecologicaltraitdata.github.io/TraitDataList/#Front_wing_width

Front_wing_surface

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	Front_wing_surface
traitName	Front_wing_surface
trait_class	morphology
broaderTerm	https://ecologicaltraitdata.github.io/TraitDataList/#Wing_size_relation
valueType	numeric
traitUnit	cm²
maxAllowedValue	999
minAllowedValue	0
factorLevels	NA
traitDescription	total surface area of the front wing
Comments
example	2.5
relationSource	refines
source	http://t-sita.cesab.org/BETSI_vizInfo.jsp?trait=Wing_surface

Front_wing_surface_max

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	Front_wing_surface_max
traitName	Front_wing_surface_max
trait_class	morphology
broaderTerm	https://ecologicaltraitdata.github.io/TraitDataList/#Front_wing_surface
valueType	numeric
traitUnit	cm²
maxAllowedValue	999
minAllowedValue	0
factorLevels	NA
traitDescription	maximum documented total surface area of the front wing
Comments
example	2.75
relationSource	inheritsfrom
source	https://ecologicaltraitdata.github.io/TraitDataList/#Front_wing_surface

Front_wing_surface_min

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	Front_wing_surface_min
traitName	Front_wing_surface_min
trait_class	morphology
broaderTerm	https://ecologicaltraitdata.github.io/TraitDataList/#Front_wing_surface
valueType	numeric
traitUnit	cm²
maxAllowedValue	999
minAllowedValue	0
factorLevels	NA
traitDescription	minimum documented total surface area of the front wing
Comments
example	2.25
relationSource	inheritsfrom
source	https://ecologicaltraitdata.github.io/TraitDataList/#Front_wing_surface

Hind_wing_length

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	Hind_wing_length
traitName	Hind_wing_length
trait_class	morphology
broaderTerm	https://ecologicaltraitdata.github.io/TraitDataList/#Wing_size_relation
valueType	numeric
traitUnit	cm
maxAllowedValue	999
minAllowedValue	0
factorLevels	NA
traitDescription	longest dimension of the hind wing
Comments
example	3
relationSource	refines
source	http://t-sita.cesab.org/BETSI_vizInfo.jsp?trait=Wing

Hind_wing_length_max

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	Hind_wing_length_max
traitName	Hind_wing_length_max
trait_class	morphology
broaderTerm	https://ecologicaltraitdata.github.io/TraitDataList/#Hind_wing_length
valueType	numeric
traitUnit	cm
maxAllowedValue	999
minAllowedValue	0
factorLevels	NA
traitDescription	maximum documented longest dimension of the hind wing
Comments
example	4
relationSource	inheritsfrom
source	https://ecologicaltraitdata.github.io/TraitDataList/#Hind_wing_length

Hind_wing_length_min

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	Hind_wing_length_min
traitName	Hind_wing_length_min
trait_class	morphology
broaderTerm	https://ecologicaltraitdata.github.io/TraitDataList/#Hind_wing_length
valueType	numeric
traitUnit	cm
maxAllowedValue	999
minAllowedValue	0
factorLevels	NA
traitDescription	minimum documented longest dimension of the hind wing
Comments
example	2
relationSource	inheritsfrom
source	https://ecologicaltraitdata.github.io/TraitDataList/#Hind_wing_length

Hind_wing_width

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	Hind_wing_width
traitName	Hind_wing_width
trait_class	morphology
broaderTerm	https://ecologicaltraitdata.github.io/TraitDataList/#Wing_size_relation
valueType	numeric
traitUnit	cm
maxAllowedValue	999
minAllowedValue	0
factorLevels	NA
traitDescription	second longest dimension of the hind wing
Comments
example	2
relationSource	refines
source	http://t-sita.cesab.org/BETSI_vizInfo.jsp?trait=Wing

Hind_wing_width_max

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	Hind_wing_width_max
traitName	Hind_wing_width_max
trait_class	morphology
broaderTerm	https://ecologicaltraitdata.github.io/TraitDataList/#Hind_wing_width
valueType	numeric
traitUnit	cm
maxAllowedValue	999
minAllowedValue	0
factorLevels	NA
traitDescription	maximum documented second longest dimension of the hind wing
Comments
example	2.5
relationSource	inheritsfrom
source	https://ecologicaltraitdata.github.io/TraitDataList/#Hind_wing_width

Hind_wing_width_min

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	Hind_wing_width_min
traitName	Hind_wing_width_min
trait_class	morphology
broaderTerm	https://ecologicaltraitdata.github.io/TraitDataList/#Hind_wing_width
valueType	numeric
traitUnit	cm
maxAllowedValue	999
minAllowedValue	0
factorLevels	NA
traitDescription	minimum documented second longest dimension of the hind wing
Comments
example	1.5
relationSource	inheritsfrom
source	https://ecologicaltraitdata.github.io/TraitDataList/#Hind_wing_width

Hind_wing_surface

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	Hind_wing_surface
traitName	Hind_wing_surface
trait_class	morphology
broaderTerm	https://ecologicaltraitdata.github.io/TraitDataList/#Wing_size_relation
valueType	numeric
traitUnit	cm²
maxAllowedValue	999
minAllowedValue	0
factorLevels	NA
traitDescription	total surface area of the hind wing
Comments
example	2.5
relationSource	refines
source	http://t-sita.cesab.org/BETSI_vizInfo.jsp?trait=Wing_surface

Hind_wing_surface_max

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	Hind_wing_surface_max
traitName	Hind_wing_surface_max
trait_class	morphology
broaderTerm	https://ecologicaltraitdata.github.io/TraitDataList/#Hind_wing_surface
valueType	numeric
traitUnit	cm²
maxAllowedValue	999
minAllowedValue	0
factorLevels	NA
traitDescription	maximum documented total surface area of the hind wing
Comments
example	2.75
relationSource	inheritsfrom
source	https://ecologicaltraitdata.github.io/TraitDataList/#Hind_wing_surface

Hind_wing_surface_min

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	Hind_wing_surface_min
traitName	Hind_wing_surface_min
trait_class	morphology
broaderTerm	https://ecologicaltraitdata.github.io/TraitDataList/#Hind_wing_surface
valueType	numeric
traitUnit	cm²
maxAllowedValue	999
minAllowedValue	0
factorLevels	NA
traitDescription	minimum documented total surface area of the hind wing
Comments
example	2.25
relationSource	inheritsfrom
source	https://ecologicaltraitdata.github.io/TraitDataList/#Hind_wing_surface

Wing_appearance_Females

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	Wing_appearance_Females
traitName	Wing_appearance_Females
trait_class	morphology
broaderTerm	https://ecologicaltraitdata.github.io/TraitDataList/#Wing_appearance
valueType	factor
traitUnit	NA
maxAllowedValue	NA
minAllowedValue	NA
factorLevels	apter/hypopter/submacropter/macropter/brachypter
traitDescription	type of membranous wings in females
Comments
example	apter
relationSource	refines
source	https://ecologicaltraitdata.github.io/TraitDataList/#Wing_appearance

Wing_appearance_Males

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	Wing_appearance_Males
traitName	Wing_appearance_Males
trait_class	morphology
broaderTerm	https://ecologicaltraitdata.github.io/TraitDataList/#Wing_appearance
valueType	factor
traitUnit	NA
maxAllowedValue	NA
minAllowedValue	NA
factorLevels	apter/hypopter/submacropter/macropter/brachypter
traitDescription	type of membranous wings in males
Comments
example	apter
relationSource	refines
source	https://ecologicaltraitdata.github.io/TraitDataList/#Wing_appearance

Front_wing_length

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	Front_wing_length
traitName	Front_wing_length
trait_class	morphology
broaderTerm	https://ecologicaltraitdata.github.io/TraitDataList/#Wing_size_relation
valueType	numeric
traitUnit	cm
maxAllowedValue	999
minAllowedValue	0
factorLevels	NA
traitDescription	longest dimension of the front wing
Comments
example	3
relationSource	refines
source	http://t-sita.cesab.org/BETSI_vizInfo.jsp?trait=Wing

Front_wing_length_max

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	Front_wing_length_max
traitName	Front_wing_length_max
trait_class	morphology
broaderTerm	https://ecologicaltraitdata.github.io/TraitDataList/#Front_wing_length
valueType	numeric
traitUnit	cm
maxAllowedValue	999
minAllowedValue	0
factorLevels	NA
traitDescription	maximum documented longest dimension of the front wing
Comments
example	4
relationSource	inheritsfrom
source	https://ecologicaltraitdata.github.io/TraitDataList/#Front_wing_length

Front_wing_length_min

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	Front_wing_length_min
traitName	Front_wing_length_min
trait_class	morphology
broaderTerm	https://ecologicaltraitdata.github.io/TraitDataList/#Front_wing_length
valueType	numeric
traitUnit	cm
maxAllowedValue	999
minAllowedValue	0
factorLevels	NA
traitDescription	minimum documented longest dimension of the front wing
Comments
example	2
relationSource	inheritsfrom
source	https://ecologicaltraitdata.github.io/TraitDataList/#Front_wing_length

Elytra_length

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	Elytra_length
traitName	Elytra_length
trait_class	morphology
broaderTerm	https://ecologicaltraitdata.github.io/TraitDataList/#Wing_appearance
valueType	numeric
traitUnit	cm
maxAllowedValue	999
minAllowedValue	0
factorLevels	NA
traitDescription	longest dimension of the elytra
Comments
example	3
relationSource	refines
source	http://t-sita.cesab.org/BETSI_vizInfo.jsp?trait=Wing

Elytra_length_max

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	Elytra_length_max
traitName	Elytra_length_max
trait_class	morphology
broaderTerm	https://ecologicaltraitdata.github.io/TraitDataList/#Elytra_length
valueType	numeric
traitUnit	cm
maxAllowedValue	999
minAllowedValue	0
factorLevels	NA
traitDescription	maximum documented longest dimension of the elytra
Comments
example	4
relationSource	inheritsfrom
source	https://ecologicaltraitdata.github.io/TraitDataList/#Elytra_length

Elytra_length_min

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	Elytra_length_min
traitName	Elytra_length_min
trait_class	morphology
broaderTerm	https://ecologicaltraitdata.github.io/TraitDataList/#Elytra_length
valueType	numeric
traitUnit	cm
maxAllowedValue	999
minAllowedValue	0
factorLevels	NA
traitDescription	minimum documented longest dimension of the elytra
Comments
example	2
relationSource	inheritsfrom
source	https://ecologicaltraitdata.github.io/TraitDataList/#Elytra_length

Elytra_width

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	Elytra_width
traitName	Elytra_width
trait_class	morphology
broaderTerm	https://ecologicaltraitdata.github.io/TraitDataList/#Wing_appearance
valueType	numeric
traitUnit	cm
maxAllowedValue	999
minAllowedValue	0
factorLevels	NA
traitDescription	second longest dimension of the elytra
Comments
example	2
relationSource	refines
source	http://t-sita.cesab.org/BETSI_vizInfo.jsp?trait=Wing

Elytra_width_max

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	Elytra_width_max
traitName	Elytra_width_max
trait_class	morphology
broaderTerm	https://ecologicaltraitdata.github.io/TraitDataList/#Elytra_width
valueType	numeric
traitUnit	cm
maxAllowedValue	999
minAllowedValue	0
factorLevels	NA
traitDescription	maximum documented second longest dimension of the elytra
Comments
example	2.5
relationSource	inheritsfrom
source	https://ecologicaltraitdata.github.io/TraitDataList/#Elytra_width

Elytra_width_min

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	Elytra_width_min
traitName	Elytra_width_min
trait_class	morphology
broaderTerm	https://ecologicaltraitdata.github.io/TraitDataList/#Elytra_width
valueType	numeric
traitUnit	cm
maxAllowedValue	999
minAllowedValue	0
factorLevels	NA
traitDescription	minimum documented second longest dimension of the elytra
Comments
example	1.5
relationSource	inheritsfrom
source	https://ecologicaltraitdata.github.io/TraitDataList/#Elytra_width

Elytra_surface

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	Elytra_surface
traitName	Elytra_surface
trait_class	morphology
broaderTerm	https://ecologicaltraitdata.github.io/TraitDataList/#Wing_appearance
valueType	numeric
traitUnit	cm²
maxAllowedValue	999
minAllowedValue	0
factorLevels	NA
traitDescription	total surface area of the elytra
Comments
example	2.5
relationSource	refines
source	http://t-sita.cesab.org/BETSI_vizInfo.jsp?trait=Wing_surface

Elytra_surface_max

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	Elytra_surface_max
traitName	Elytra_surface_max
trait_class	morphology
broaderTerm	https://ecologicaltraitdata.github.io/TraitDataList/#Elytra_surface
valueType	numeric
traitUnit	cm²
maxAllowedValue	999
minAllowedValue	0
factorLevels	NA
traitDescription	maximum documented total surface area of the elytra
Comments
example	2.75
relationSource	inheritsfrom
source	https://ecologicaltraitdata.github.io/TraitDataList/#Elytra_surface

Elytra_surface_min

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	Elytra_surface_min
traitName	Elytra_surface_min
trait_class	morphology
broaderTerm	https://ecologicaltraitdata.github.io/TraitDataList/#Elytra_surface
valueType	numeric
traitUnit	cm²
maxAllowedValue	999
minAllowedValue	0
factorLevels	NA
traitDescription	minimum documented total surface area of the elytra
Comments
example	2.25
relationSource	inheritsfrom
source	https://ecologicaltraitdata.github.io/TraitDataList/#Elytra_surface

Radial_nervure_length

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	Radial_nervure_length
traitName	Radial_nervure_length
trait_class	morphology
broaderTerm	https://ecologicaltraitdata.github.io/TraitDataList/#Wing_appearance
valueType	numeric
traitUnit	cm
maxAllowedValue	999
minAllowedValue	0
factorLevels	NA
traitDescription	length of the radial nervure of a wing
Comments
example	2.5
relationSource	equals
source	http://t-sita.cesab.org/BETSI_vizInfo.jsp?trait=Radial_nervure_length

Radial_nervure_length_max

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	Radial_nervure_length_max
traitName	Radial_nervure_length_max
trait_class	morphology
broaderTerm	https://ecologicaltraitdata.github.io/TraitDataList/#Radial_nervure_length
valueType	numeric
traitUnit	cm
maxAllowedValue	999
minAllowedValue	0
factorLevels	NA
traitDescription	maximum documented length of the radial nervure of a wing
Comments
example	3
relationSource	inheritsfrom
source	https://ecologicaltraitdata.github.io/TraitDataList/#Radial_nervure_length

Radial_nervure_length_min

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	Radial_nervure_length_min
traitName	Radial_nervure_length_min
trait_class	morphology
broaderTerm	https://ecologicaltraitdata.github.io/TraitDataList/#Radial_nervure_length
valueType	numeric
traitUnit	cm
maxAllowedValue	999
minAllowedValue	0
factorLevels	NA
traitDescription	minimum documented length of the radial nervure of a wing
Comments
example	2
relationSource	inheritsfrom
source	https://ecologicaltraitdata.github.io/TraitDataList/#Radial_nervure_length

Median_nervure_length

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	Median_nervure_length
traitName	Median_nervure_length
trait_class	morphology
broaderTerm	https://ecologicaltraitdata.github.io/TraitDataList/#Wing_appearance
valueType	numeric
traitUnit	cm
maxAllowedValue	999
minAllowedValue	0
factorLevels	NA
traitDescription	length of the median nervure of a wing
Comments
example	2
relationSource	equals
source	http://t-sita.cesab.org/BETSI_vizInfo.jsp?trait=Median_nervure_length

Median_nervure_length_max

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	Median_nervure_length_max
traitName	Median_nervure_length_max
trait_class	morphology
broaderTerm	https://ecologicaltraitdata.github.io/TraitDataList/#Median_nervure_length
valueType	numeric
traitUnit	cm
maxAllowedValue	999
minAllowedValue	0
factorLevels	NA
traitDescription	maximum documented length of the median nervure of a wing
Comments
example	2
relationSource	inheritsfrom
source	https://ecologicaltraitdata.github.io/TraitDataList/#Median_nervure_length

Median_nervure_length_min

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	Median_nervure_length_min
traitName	Median_nervure_length_min
trait_class	morphology
broaderTerm	https://ecologicaltraitdata.github.io/TraitDataList/#Median_nervure_length
valueType	numeric
traitUnit	cm
maxAllowedValue	999
minAllowedValue	0
factorLevels	NA
traitDescription	minimum documented length of the median nervure of a wing
Comments
example	2
relationSource	inheritsfrom
source	https://ecologicaltraitdata.github.io/TraitDataList/#Median_nervure_length

Front_wing_span

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	Front_wing_span
traitName	Front_wing_span
trait_class	morphology
broaderTerm	https://ecologicaltraitdata.github.io/TraitDataList/#Locomotion_morphology
valueType	numeric
traitUnit	cm
maxAllowedValue	999
minAllowedValue	0
factorLevels	NA
traitDescription	distance between the outer tips of the front wings when in flight
Comments
example	12
relationSource
source

Front_wing_span_max

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	Front_wing_span_max
traitName	Front_wing_span_max
trait_class	morphology
broaderTerm	https://ecologicaltraitdata.github.io/TraitDataList/#Front_wing_span
valueType	numeric
traitUnit	cm
maxAllowedValue	999
minAllowedValue	0
factorLevels	NA
traitDescription	maximum documented distance between the outer tips of the front wings when in flight
Comments
example	15
relationSource	inheritsfrom
source	https://ecologicaltraitdata.github.io/TraitDataList/#Front_wing_span

Front_wing_span_min

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	Front_wing_span_min
traitName	Front_wing_span_min
trait_class	morphology
broaderTerm	https://ecologicaltraitdata.github.io/TraitDataList/#Front_wing_span
valueType	numeric
traitUnit	cm
maxAllowedValue	999
minAllowedValue	0
factorLevels	NA
traitDescription	minimum documented distance between the outer tips of the front wings when in flight
Comments
example	10
relationSource	inheritsfrom
source	https://ecologicaltraitdata.github.io/TraitDataList/#Front_wing_span

Hind_wing_span

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	Hind_wing_span
traitName	Hind_wing_span
trait_class	morphology
broaderTerm	https://ecologicaltraitdata.github.io/TraitDataList/#Locomotion_morphology
valueType	numeric
traitUnit	cm
maxAllowedValue	999
minAllowedValue	0
factorLevels	NA
traitDescription	distance between the outer tips of the hind wings when in flight
Comments
example	12
relationSource
source

Hind_wing_span_max

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	Hind_wing_span_max
traitName	Hind_wing_span_max
trait_class	morphology
broaderTerm	https://ecologicaltraitdata.github.io/TraitDataList/#Hind_wing_span
valueType	numeric
traitUnit	cm
maxAllowedValue	999
minAllowedValue	0
factorLevels	NA
traitDescription	maximum documented distance between the outer tips of the hind wings when in flight
Comments
example	15
relationSource	inheritsfrom
source	https://ecologicaltraitdata.github.io/TraitDataList/#Hind_wing_span

Hind_wing_span_min

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	Hind_wing_span_min
traitName	Hind_wing_span_min
trait_class	morphology
broaderTerm	https://ecologicaltraitdata.github.io/TraitDataList/#Hind_wing_span
valueType	numeric
traitUnit	cm
maxAllowedValue	999
minAllowedValue	0
factorLevels	NA
traitDescription	minimum documented distance between the outer tips of the hind wings when in flight
Comments
example	10
relationSource	inheritsfrom
source	https://ecologicaltraitdata.github.io/TraitDataList/#Hind_wing_span

Saltatory_apparatus

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	Saltatory_apparatus
traitName	Saltatory_apparatus
trait_class	morphology
broaderTerm	https://ecologicaltraitdata.github.io/TraitDataList/#Locomotion_morphology
valueType	character
traitUnit	NA
maxAllowedValue	NA
minAllowedValue	NA
factorLevels	NA
traitDescription	any morphological structure which serves a specialized function for jumping
Comments
example	NA
relationSource	equals
source	http://t-sita.cesab.org/BETSI_vizInfo.jsp?trait=Saltatory_apparatus

Saltatory_apparatus_length

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	Saltatory_apparatus_length
traitName	Saltatory_apparatus_length
trait_class	morphology
broaderTerm	https://ecologicaltraitdata.github.io/TraitDataList/#Saltatory_apparatus
valueType	numeric
traitUnit	cm
maxAllowedValue	999
minAllowedValue	0
factorLevels	NA
traitDescription	length of the saltatory apparatus
Comments
example	3
relationSource	equals
source	http://t-sita.cesab.org/BETSI_vizInfo.jsp?trait=Saltatory_apparatus_length

Saltatory_apparatus_length_max

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	Saltatory_apparatus_length_max
traitName	Saltatory_apparatus_length_max
trait_class	morphology
broaderTerm	https://ecologicaltraitdata.github.io/TraitDataList/#Saltatory_apparatus_length
valueType	numeric
traitUnit	cm
maxAllowedValue	999
minAllowedValue	0
factorLevels	NA
traitDescription	maximum documented length of the saltatory apparatus
Comments
example	4
relationSource	inheritsfrom
source	https://ecologicaltraitdata.github.io/TraitDataList/#Saltatory_apparatus_length

Saltatory_apparatus_length_min

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	Saltatory_apparatus_length_min
traitName	Saltatory_apparatus_length_min
trait_class	morphology
broaderTerm	https://ecologicaltraitdata.github.io/TraitDataList/#Saltatory_apparatus_length
valueType	numeric
traitUnit	cm
maxAllowedValue	999
minAllowedValue	0
factorLevels	NA
traitDescription	minimum documented length of the saltatory apparatus
Comments
example	2
relationSource	inheritsfrom
source	https://ecologicaltraitdata.github.io/TraitDataList/#Saltatory_apparatus_length

Leg_body_ratio

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	Leg_body_ratio
traitName	Leg_body_ratio
trait_class	morphology
broaderTerm	https://ecologicaltraitdata.github.io/TraitDataList/#morphology
valueType	numeric
traitUnit	ratio
maxAllowedValue	999
minAllowedValue	0
factorLevels	NA
traitDescription	ration between leg length and body lengt
Comments
example	2
relationSource
source

Leg_body_ratio_max

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	Leg_body_ratio_max
traitName	Leg_body_ratio_max
trait_class	morphology
broaderTerm	https://ecologicaltraitdata.github.io/TraitDataList/#Leg_body_ratio
valueType	numeric
traitUnit	ratio
maxAllowedValue	999
minAllowedValue	0
factorLevels	NA
traitDescription	maximum documented ration between leg length and body lengt
Comments
example	2.25
relationSource	inheritsfrom
source	https://ecologicaltraitdata.github.io/TraitDataList/#Leg_body_ratio

Leg_body_ratio_min

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	Leg_body_ratio_min
traitName	Leg_body_ratio_min
trait_class	morphology
broaderTerm	https://ecologicaltraitdata.github.io/TraitDataList/#Leg_body_ratio
valueType	numeric
traitUnit	ratio
maxAllowedValue	999
minAllowedValue	0
factorLevels	NA
traitDescription	minimum documented ration between leg length and body lengt
Comments
example	1.75
relationSource	inheritsfrom
source	https://ecologicaltraitdata.github.io/TraitDataList/#Leg_body_ratio

Protection_morphology

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	Protection_morphology
traitName	Protection_morphology
trait_class	morphology
broaderTerm	https://ecologicaltraitdata.github.io/TraitDataList/#morphology
valueType	character
traitUnit	NA
maxAllowedValue	NA
minAllowedValue	NA
factorLevels	NA
traitDescription	any morphological structure which serves a specialized function for protection against biotic and abiotic threats
Comments
example	NA
relationSource	equals
source	http://t-sita.cesab.org/BETSI_vizInfo.jsp?trait=Protection

Body_color

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	Body_color
traitName	Body_color
trait_class	morphology
broaderTerm	https://ecologicaltraitdata.github.io/TraitDataList/#Protection_morphology
valueType	character
traitUnit	NA
maxAllowedValue	NA
minAllowedValue	NA
factorLevels	NA
traitDescription	color of the main body
Comments
example	NA
relationSource	equals
source	http://t-sita.cesab.org/BETSI_vizInfo.jsp?trait=Body_color

Body_color_gradient

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	Body_color_gradient
traitName	Body_color_gradient
trait_class	morphology
broaderTerm	https://ecologicaltraitdata.github.io/TraitDataList/#Body_color
valueType	character
traitUnit	NA
maxAllowedValue	NA
minAllowedValue	NA
factorLevels	NA
traitDescription	color gradient of the main body
Comments
example	NA
relationSource	equals
source	http://t-sita.cesab.org/BETSI_vizInfo.jsp?trait=Body_color_gradient

Body_color_pattern

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	Body_color_pattern
traitName	Body_color_pattern
trait_class	morphology
broaderTerm	https://ecologicaltraitdata.github.io/TraitDataList/#Body_color
valueType	character
traitUnit	NA
maxAllowedValue	NA
minAllowedValue	NA
factorLevels	NA
traitDescription	color pattern of the main body
Comments
example	NA
relationSource	equals
source	http://t-sita.cesab.org/BETSI_vizInfo.jsp?trait=Body_color_pattern

Repellent_glands

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	Repellent_glands
traitName	Repellent_glands
trait_class	morphology
broaderTerm	https://ecologicaltraitdata.github.io/TraitDataList/#Protection_morphology
valueType	logical
traitUnit	NA
maxAllowedValue	NA
minAllowedValue	NA
factorLevels	NA
traitDescription	presence of glands secreting noxious liquids or vapors to repel antagonists
Comments	includes synonyms repugnatorial glands and repulsive glands
example	TRUE
relationSource	equals
source	http://t-sita.cesab.org/BETSI_vizInfo.jsp?trait=Repellent_glands

Sclerotization

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	Sclerotization
traitName	Sclerotization
trait_class	morphology
broaderTerm	https://ecologicaltraitdata.github.io/TraitDataList/#Protection_morphology
valueType	logical
traitUnit	NA
maxAllowedValue	NA
minAllowedValue	NA
factorLevels	NA
traitDescription	presence of hardened body surface by deposition of sclerotin or other substances in the external covering of the physical structure of organisms
Comments	includes synonyms cuticle or integument sclerotization/sclerification
example	TRUE
relationSource	equals
source	http://t-sita.cesab.org/BETSI_vizInfo.jsp?trait=Repellent_glands

Detoxication_strategy

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	Detoxication_strategy
traitName	Detoxication_strategy
trait_class	morphology
broaderTerm	https://ecologicaltraitdata.github.io/TraitDataList/#Protection_morphology
valueType	character
traitUnit	NA
maxAllowedValue	NA
minAllowedValue	NA
factorLevels	NA
traitDescription	any morphological structure or ability which serves a specialized function to eliminate or neutralize a toxin
Comments
example	NA
relationSource	equals
source	http://t-sita.cesab.org/BETSI_vizInfo.jsp?trait=Detoxication_strategies

Water_regulation_morphology

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	Water_regulation_morphology
traitName	Water_regulation_morphology
trait_class	morphology
broaderTerm	https://ecologicaltraitdata.github.io/TraitDataList/#Protection_morphology
valueType	character
traitUnit	NA
maxAllowedValue	NA
minAllowedValue	NA
factorLevels	NA
traitDescription	any morphological structure which ensures an hydric status of the animal
Comments
example	NA
relationSource	refines
source	http://t-sita.cesab.org/BETSI_vizInfo.jsp?trait=waterregulation%20strategies

Traits describing physiology

Water_regulation_physiology

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	Water_regulation_physiology
traitName	Water_regulation_physiology
trait_class	physiology
broaderTerm	https://ecologicaltraitdata.github.io/TraitDataList/#physiology
valueType	character
traitUnit	NA
maxAllowedValue	NA
minAllowedValue	NA
factorLevels	NA
traitDescription	any physiological mechanism which ensures an hydric status of the animal
Comments
example	NA
relationSource	refines
source	http://t-sita.cesab.org/BETSI_vizInfo.jsp?trait=waterregulation%20strategies

Thermal_regulation_physiology

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	Thermal_regulation_physiology
traitName	Thermal_regulation_physiology
trait_class	physiology
broaderTerm	https://ecologicaltraitdata.github.io/TraitDataList/#physiology
valueType	character
traitUnit	NA
maxAllowedValue	NA
minAllowedValue	NA
factorLevels	NA
traitDescription	any physiological mechanism which keeps the body temperature within certain boundaries
Comments
example	NA
relationSource	refines
source	http://t-sita.cesab.org/BETSI_vizInfo.jsp?trait=Thermoregulation_strategies

Water_loss_rate

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	Water_loss_rate
traitName	Water_loss_rate
trait_class	physiology
broaderTerm	https://ecologicaltraitdata.github.io/TraitDataList/#physiology
valueType	numeric
traitUnit	mg/h
maxAllowedValue	999
minAllowedValue	0
factorLevels	NA
traitDescription	rate of water loss at ambient temperature
Comments
example	0.5
relationSource
source

Larval_inundation_tolerance

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	Larval_inundation_tolerance
traitName	Larval_inundation_tolerance
trait_class	physiology
broaderTerm	https://ecologicaltraitdata.github.io/TraitDataList/#physiology
valueType	numeric
traitUnit	hour
maxAllowedValue	9999
minAllowedValue	0
factorLevels	NA
traitDescription	maximum time under water after which larvae resume development
Comments
example	12
relationSource
source

Venom_excretion

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	Venom_excretion
traitName	Venom_excretion
trait_class	physiology
broaderTerm	https://ecologicaltraitdata.github.io/TraitDataList/#physiology
valueType	character
traitUnit	NA
maxAllowedValue	NA
minAllowedValue	NA
factorLevels	NA
traitDescription	physiological mechanism by which venom is secreted
Comments
example	NA
relationSource	equals
source	http://t-sita.cesab.org/BETSI_vizInfo.jsp?trait=Venom_excretion

Regeneration_strategy

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	Regeneration_strategy
traitName	Regeneration_strategy
trait_class	physiology
broaderTerm	https://ecologicaltraitdata.github.io/TraitDataList/#physiology
valueType	character
traitUnit	NA
maxAllowedValue	NA
minAllowedValue	NA
factorLevels	NA
traitDescription	any physiological mechanism which allows replacement of body parts
Comments
example	NA
relationSource	equals
source	http://t-sita.cesab.org/BETSI_vizInfo.jsp?trait=Regeneration_strategies

Traits relating to pollination

Flower_color_preference

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	Flower_color_preference
traitName	Flower_color_preference
trait_class	pollination
broaderTerm	https://ecologicaltraitdata.github.io/TraitDataList/#pollination
valueType	character
traitUnit	NA
maxAllowedValue	NA
minAllowedValue	NA
factorLevels	NA
traitDescription	color or set of colors which is preferably approached
Comments
example	NA
relationSource
source

Flower_visitor

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	Flower_visitor
traitName	Flower_visitor
trait_class	pollination
broaderTerm	https://ecologicaltraitdata.github.io/TraitDataList/#pollination
valueType	logical
traitUnit	NA
maxAllowedValue	NA
minAllowedValue	NA
factorLevels	NA
traitDescription	individuals visit flowers to collect pollen and/or nectar
Comments
example	TRUE
relationSource
source

Flower_specialisation

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	Flower_specialisation
traitName	Flower_specialisation
trait_class	pollination
broaderTerm	https://ecologicaltraitdata.github.io/TraitDataList/#Flower_visitor
valueType	character
traitUnit	NA
maxAllowedValue	NA
minAllowedValue	NA
factorLevels	NA
traitDescription	type of flower which is preferably approached
Comments
example	NA
relationSource
source

Flower_feeding

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	Flower_feeding
traitName	Flower_feeding
trait_class	pollination
broaderTerm	https://ecologicaltraitdata.github.io/TraitDataList/#Flower_visitor
valueType	factor
traitUnit	NA
maxAllowedValue	NA
minAllowedValue	NA
factorLevels	nectar/pollen/both
traitDescription	nutrient source selected by flower visitors
Comments
example	nectar
relationSource
source

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Ecological Traitdata Standard by Nadja K. Simons, Florian D. Schneider, Caterina Penone, Andreas Ostrowski is licensed under a Creative Commons Attribution 4.0 International License.